GAMETOGBNESIS OF SACCOCIRRUS » 



have become drawn out to form a comet-tail body attached 

 to the spermatid nucleus. The latter has begun to undergo 

 the usual changes which may be seen in figs. 9, 10, 11, 12, and 

 13. In PI. '2, fig. 16, the tail has further elongated, the yolk- 

 spherules and Golgi elements are drifting down the length of 

 the tail, while the nucleus is losing its reticular arrangement. 

 This lengthening process now goes on till the fully-formed 

 elongate Saccocirrus spermatozoon is produced. 



Owing to the small size of the cells, and possibly to the 

 unsuitability of the material, no satisfactory description can 

 be given of the formation of the acrosome. This can be seen 

 in PI. 2, fig. 13, at as. In figs. 9 and 10, at gx, were bodies 

 which might have some connexion with the formation of the 

 acrosome, but of this 1 am unable to speak with certainty. 



(/) The Fate of the Golgi Apparatus during 

 S p e r m a t e 1 e s i s . 



If one examines a bundle of ripe or ripening spermatozoa 

 in material prepared by the Mann-Kopsch method, it will be 

 found that near the end of the tail of each sperm are rounded 

 or angular bodies which are stained by the osmic acid : in 

 PI. 2, fig. 17, is drawn such a bundle of sperms, the bodies 

 being marked gax ; at a higher magnification, as in fig. 15, 

 the bodies are seen to be very like the Golgi body drawn in 

 the spermatogonium in PI. 1, fig. 7, at ga. I cannot say for 

 certain whether these bodies are derived from the original 

 Golgi rods depicted in PI. 2, figs. 8-13, but I should think that 

 they are so derived, and that they have undergone some 

 change during late spermateleosis. If these bodies are an 

 integral part of the spermatozoon and not the degeneration 

 products of the spermateleosis, one might expect to find 

 some sign of them in the spermatozoa within the female : 

 in PI. 2, fig. 1 8, is section of the receptaculum se minis 

 of a female, and drawn to the same scale as fig. 1 7 above. The 

 same bodies are to be seen at the tails of the ripe sperma- 

 tozoa. 



