CALMA GLAUCOIDES 445 



and indented by the pericardium {p.c.s.) and anteriorly to that 

 by the ootype (n.g.s.). Thus two chambers are connected by 

 a narrow tube, but neither their histological structure nor their 

 function justifies their being regarded as other than mechanically 

 separated portions of a continuous food reservoir. The lining- 

 cells of the gastric sac are low and fiat even when it is more or 

 less empty, while the cells of the ceratal diverticula are very 

 large and extensively vacuolated during active digestion. 

 A comparison of these hepato-pancreatic cells in the active 

 state (fig. 5, etc.), and the tenuous squames that line the full 

 stomach (fig. 8), strongly suggests that the former are responsible 

 for the bulk of the digestive juices in Calma. The fish embryos 

 (fig. 8, s.e.), whether very young or considerably developed 

 when eaten, are, however, partly disintegrated in situ in 

 the stomach, probably by enzymes delivered from the ceratal 

 glands (fig. 8, d.l.), which at this stage are uninvaded by the 

 food. Later the stomach contains a semifluid mass in which 

 lenses of eyes (fig. 8, /.) and scattered lumps of undigested 

 nuclei are the only remaining evidences of the nature of the 

 food. During further digestion this thick fluid is continuously 

 delivered into the cerata, where it undergoes solution (fig. 9, 

 d.l.). An animal fixed at this stage is difficult to cut on account 

 of the extremely hard consistency of the precipitated proteins. 

 The gland-cells of the ceratal outgrowths (fig. 5, d.c.) project 

 deeply into the food ; no evidence of ingestion could be found, 

 though fine brown granules similar to the eventual residuum 

 in the whole system accumulate in them and are extruded into 

 the lumen. 



There is no trace of anus or intestine. The small amount 

 of undigested matter remains as a dark- brown core (fig. 9, d.) 

 throughout the alimentary system, so that the shape of this 

 system can be made out in a fasting animal by clearing alone. 

 On account of a certain amount of compression of this faecal 

 residue during the fast there is no admixture with a subsequent 

 meal. In connexion with the digestive system must be men- 

 tioned certain special connective-tissue cells (figs. 8 and 9, c.s.) 

 of the cerata which differ widely from those of the rest of the 



NO= 263 li 



