1064 
In crossing common blue flax AABBCC with the narrow-petalled 
white AACC, which differ in breadth and in colour, a complete 
connection was observed between these two characters. All 
blue descendants had the broadness-type of the common blue 
flax, all white ones were of the narrow-type like the white P-varie- 
ty. This is now also quite intelligible. It was shown above that 
the narrow-petalled white flax has the same factors for broadness as 
the common blue. All offspring from the crossing of these two 
varieties will therefore have the same factors for broadness. Now all 
the blue descendants possess, in addition to these factors for broad- 
ness also both the factors B and C, the white ones however have, 
besides the factors for broadness, only C. In all the blue therefore 
the factors for broadness are undisturbed, they will in consequence 
all have the breadth of the blue P-variety. In the white ones on 
the other hand the factors for broadness are inhibited, they will be 
narrow like the white P-variety. It follows that in this case also 
the interdependence between breadth and colour is other than can 
be deduced off hand from this crossing. The blue colour and the 
greater breadth of the common blue flax are not caused by the 
presence of a single factor or group of “factors, in the absence of 
which the colour is white and the breadth smaller, but the relation- 
ship is a different one. The factor B which is present in the common 
blue ‘flax, and absent from the narrow-petalled white, is not at the 
same time a factor for both, colour and broadness. In the two varie- 
ties there are present broadness-factors other than B and they are 
the same for both; but in the narrow-petalled white they are inhi- 
bited by C, in the common variety they are unaffected owing to 
the presence of B and C together. 
Further the more complicated phenomena which were observed 
in the crossing between Egyptian AABBCC and the narrow-petalled 
white flax AACC, can now be completely explained. The difference 
between these two varieties is, so far as the factors for broadness are 
concerned, exactly the same as that between Egyptian and common 
blue and as that between Egyptian and common white flax. For 
the narrow-petalled white flax has the same factors for broadness 
as the common blue and the common white. Now since the broadness- 
factors follow the segregation law quite independently of the presence 
or absence of the factors B and C, the offspring will agree perfectly, 
as regards presence of broadness-factors, with the offspring from the 
crossings of the Egyptian flax with the common blue or the common 
white. 
If no other causes were acting the individuals of /, would together 
