16 BROOKLYN BOTANIC GARDEN MEMOIRS 
to note that the forms with parthenogenetic asci, generalized or not, 
were not endowed with potentialities of progress, nor with the evolu- 
tion of any important lines. They have made practically little progress 
and are few in number. On the other hand, those forms with sexually 
produced asci, even though the sexuality be of a very greatly reduced 
type, were endowed with great potentialities as evidenced by the large 
group of Euascomycetes with high specialization, and great divergence 
of character in several different series. 
TECHNICAL DESCRIPTION 
Endogone sphagnophila’ n. sp. Plants (zygocarps), 2-4 mm. in 
diameter, pulvinate, reniform, plain or subcerebriform with two to 
three low lobes or convolutions, orange yellow when mature, Peridium 
white, submembranous, tough, of interwoven coenocytic, profusely 
branched hyphae, minutely tomentose or downy from free, terminal, 
very slender branchlets, 4-5 wat base, 1 worlessat the tips. Mycelium 
of the zygocarp 10-15 » in diameter, coenocytic, stout, non-septate, 
branched in a dichotomous, or trichotomous manner, or several 
branches springing from enlargements, radial, the terminal branches 
interlacing to form the peridium. Progametes equal. Gametangia 
separated from mycelium by a cross wall, equal or usually slightly 
unequal, multinucleate. ‘‘ Resting spores’’ (zygotes) formed as an 
outgrowth from the conjugation point of the gametangia, or more rarely 
from the larger one, one resting spore formed in the primary zygote 
membrane from each pair of gametangia, elliptical to oval, rarely 
irregular, with orange yellow content and a thick, white, stratified 
cartilaginous wall, 35-60 x 30-45 yw, germination unknown. 
On sphagnum in a ravine in region of Seventh Lake, Fulton Chain, 
Adirondack Mts., New York, July 1916, Aug. 1917; and in Cranesville 
moor, Western Maryland, Sept. 1917 (rarely on other mosses or on 
dead twigs). Thaxter (Bot. Gaz. 24: 12, 1897) reports it on sphagnum 
in Maine. 
Latin diagnosis. Pulvinatis, reniformibus, subcerebriformibus, aureis, 2-4 mm.; 
peridiis albidis submenbranceis, lentis, floccosis intertextis, tomentosulis; myceliis 
glebae 12-15 », radiatis, dichotomis vel trichotonis vel plurichotomis, ramulis 
terminalibus peridium formantibus; sporis orientibus ab gametangiis copulantibus, 
ellipsoideis vel ovalibus, maximis, 35-60 x 30-45 u, plasmate aureo. Hab. on sphag- 
num, Adirondack Mts., New York, and in Maine and Maryland. 
LITERATURE CITED 
Atkinson, Geo. F. Phylogeny and Relationships in the Ascomycetes. Ann. Mo. 
Bot. Garden 2: 315-376. Figs. I-9. I914. 
7 Closely related to E. ludwigiit Bucholtz, but this species is subterranean with a 
prominent germ pore in the thick wall of the zygote and the nuclei are reduced to 
two which fuse at maturity. 
