REED: SPECIALIZATION OF PARASITIC FUNGI 365 
1. Thymi-Stipae WKleb.; aecidial stage on Thymus serpyphyllum, T. 
angustifolius and T. vulgaris. 
2. Salviae-Stipae Kleb.; aecidial stage on Salvia silvestris and S. 
pratensis. 
Puccinia sessilis Schneid. This rust, which has its uredo and 
teleuto stages on Phalaris arundinacea and its aecidial stage on various 
plants of the Liliaceae, Amaryllidaceae, Araceae and Orchidaceae, has 
been broken up into a number of distinct species, based largely upon 
the choice of the aecidial host. Klebahn (77-83, 86, 87, 89-91) has 
carried out the cultural tests with the rust and recognizes the following 
relations: 
1. Puccinia Smilacearum-Digraphidis (Sopp.) Kleb. 
a. Smilacearum-Digraphidis typica Kleb.; aecidial stage on Con- 
vallaria majalis, Maianthemum bifolium, Polygonatum 
multiflorum, P. officinale, P. verticillatum and Paris quadri- 
folia. 
b. Convallariae-Digraphidis (Sopp.) Kleb.; aecidial stage on 
Convallaria majalis. 
c. Paridi-Digraphidis (Plowr.) Kleb.; aecidial stage on Paris 
quadrifolia. 
. Puccinia Allii-Phalaridis Kleb.; aecidial stage on Allium ursinum. 
3. Puccinia Orchidearum-Phalaridis Kleb.; aecidial stage on Gymna- 
denia conopea, Listera ovata, Orchis maculata, O. latifolia, 
Platanthera bifolia and P. Chlorantha. 
4. Puccinia Ari-Phalaridis (Plowr.) Kleb.; aecidial stage on Arum 
maculatum. 
5. Puccinia Schmidtiana Diet.; aecidial stage on Leucojum aestivum 
and L. vernum. 
It may be especially noted that, according to Klebahn, Puccinia 
Smilacearum-Digraphidis includes a race which occurs on a number of 
hosts and two races which occur on only one host. The hosts for the 
second two races are also hosts for the first race. It is further to be 
noted that the species recognized are also to be distinguished on the 
basis of host relations. 
Puccinia Caricis (Schum.) Rebent. Klebahn (91) indicates the 
following specialization in the Urtica-Carex rust, the races being indi- 
cated by the choice of uredo and teleuto host: 
1. Urticae-acutae on Carex acuta, C. Goodenoughti, to a less extent C. 
stricta. 
2. Urticae-hirtae on Carex hirta. 
3. Urticae-acutiformis on Carex acutiformis, C. Kochiana, to a less 
extent on C. pseudocyperus. 
4. Urticae-vesicariae on Carex vesicaria. 
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