36 PROGRESS OF MICROSCOPICAL SCIENCE. 



lective vein, and attaclied to it by irregular bands of connective tissue, 

 whicla also holds the gland in place. From this horizontal mass, four 

 vertical branches arise, and lie between and next to the partitions at 

 the base of the legs, dividing the sides of the body into compartments. 

 The posterior of these four vertical lobes accompanies the middle 

 hepatic vein from its origin from the great collective vein, and is sent 

 off opi)osite tlje insertion of the fifth pair of feet. Half-way between 

 the origin of the vein and the articulation of the foot to the body, it 

 turns at a right angle, the ends of the two other lobes passing a 

 little beyond it, and ends in a blind sac, less vertical than the others, 

 slightly ascending at the end, which lies just above the insertion of the 

 second jjair of feet. The two middle lobes are directed to the col- 

 lective vein. Each lobe is flattened out somewhat and lies close to 

 the posterior wall of the com2)artment in which it is situated, as if 

 wedged in between the wall, and the muscles between it and the 

 anterior portion of the compartment. Each lobe also accompanies 

 the bases of the first four tegumentary nerves. I could not by in- 

 jection of the gland, make out any general opening* into the cavity of 

 the body or any connection with the hej)atic or great collective vein ; 

 any attempts to inject the gland from the veins failing. The four 

 lobes certainly end in blind sacs. The lobes are irregular in form, 

 appearing as if twisted and knotted, and with sheets and bands of con- 

 nective tissue forming the sheaths of the muscles among which the 

 gland lies. Each lobe, when cut across, is oval, with a yellowish 

 interior and a small central cavity, forming, evidently, an excretory 

 duct. The gland externally is of a bright brick red. The glandular 

 mass is quite dense, though yielding. It is singular that this con- 

 spicuous gland, though it must have engaged their attention, has not 

 been noticed by Van der Hoeven, Owen, or A. Milne-Edwards in their 

 accounts of dissections of this animal. When examined under a 

 Hartnack's No. 9 immersion lens and Zcntmayer's B eye-piece, the 

 reddish external cortical portion consists of closely aggregated irre- 

 gularly rounded nucleated cells of quite unequal size, and scattered 

 about in the interstices between the cells are dark reddish masses 

 which give colour to the gland. They are very irregular in size and 

 form, and twenty hours after a portion of the parenchyma was sub- 

 mitted to microscopic examination vibrated to and fro. I am reminded 

 in the vibrating movements of these bodies, of Siebold'sj description of 

 similar bodies in the renal organs of the Lamellibranchs, i. e. the gland 

 of Bojanus. He says in a footnote, p. 214, | ' If the walls of these 

 organs are prepared in any way for microscopic examination, a part 

 of their parencliyma separates into a vesiculo-granular mass, the 

 particles of which have a very lively dancing motion. The motions 

 are due to portions of ciliated epithelium adhering to the cells and 

 granules.' " 



" In other portions of the outer reddish part of the gland, where 



* Leydig (' Naturgeschichte der Daphnideii ') states that seveml anatomi.sts, 

 after laborious attemptB, have faihd to fiud Ihe opening to the green gland in 

 any crustacean. 



t 'Anatomy of the Invertebrates." j Burnett's Translation. 



