PRESENT POSITION OF PALEOZOIC BOTANY—SCOTT. 395 
erse the inner layers of the integument. The upper free part of the 
latter has a complex chambered structure; there are usually nine 
chambers, each of which receives one of the integumental bundles. 
The outer layer of the integument has a columnar structure. The 
free apex of the nucellus is prolonged upward through the micropyle, 
protruding somewhat beyond it, as an open tube. (Fig. 10.) “As in 
recent Cycads and in Ginkgo, the apex of the nucellus contains the 
pollen chamber, which here has a peculiar form, for the middle of the 
chamber is occupied by a solid column of tissue, reducing the actual 
‘avity to an annular channel in which the pollen grains are found. 
(See pl. 1 and fig. 10.) Within the body of the nucellus is the mem- 
brane of the megaspore or embryo sac. The seed was thus of com- 
Fic. 10.—Lagenostoma Lomazi. Apex of seed in median longitudinal section through 
micropyle. ot, outer part of testa; 7, palisade layer; it, inner part of testa; s, cavity 
between testa and nucellus; 0, orifice of pollen-chamber, pe; cc, central column; pg, 
pollen-grains ; pl, nucellus; mg, megaspore. »% about 50. After Oliver. 
plex organization and shows that Lyginodendron, in spite of its sur- 
viving Fern-like characters, had definitely attained the rank of a 
typical Spermophyte. 
The structure of the pedicel indicates that the seed was borne on a 
foliar organ. The evidence of other species leaves no doubt that the 
sporophylls were modified fronds or pinne of compound form, chiefly 
differing from the sterile foliage in the suppression of the laminz of 
the leaflets. - 
The stellate lobed indusia or cupules belonging to Calymmatotheca 
Strangeri, a species closely allied to Sphenopteris Héninghausii, is 
shown in fig. 11, drawn from. Stur’s original specimen. Another 
related type, Lagenostoma Sinclairi (fig. 12), representing both 
seeds and cupule, has been described by Arber. 
