254 TRANSACTIONS OF THE CANADIAN INSTITUTE. [Vov. VIII. 
rence, one having to examine many specimens before seeing an example, 
such as is represented in Fig. 11. The leucocyte, which is of the poly- 
morphonuclear variety is surrounded by fat, which forms a fine mesh- 
work, and yet contains none itself. It is perfectly formed and shows no 
signs whatever of disintegration. In all the preparations from the intes- 
tine of the guinea pig I have not seen half a dozen leucocytes in the chyle 
vessel, nor have I seen a leucocyte penetrating the wall, facts which should 
tell against this view. I have, moreover, never seen large lymphoid leuco- 
cytes in the lacteal,and these are the cells which, undoubtedly, pick up fat. 
Then too the fewness of leucocytes relative to the amount of fat to be 
transferred must always cast grave doubt as to the efficiency of their 
action in this respect. Finally, as I have already pointed out, one can 
frequently trace the fat as minute streams going through the parenchyma 
into the central vessel, and this without the intervention of leucocytes of 
any description. (Figs. 8 and 19). 
VIII.—FarT IN THE STRIATED BORDER. 
And now I come to the discussion of the most interesting part of my 
investigation. With the exception of Kischensky, all observers are agreed 
that fat has not been found in the striated border of the epithelium. From 
an histological standpoint, this fact has been used as a proof that fat enters 
the cells in some soluble form, for example, as a soap, and is later recon- 
verted into a neutral fat through some action of the epithelium, all observ- 
ers, further, regarding the fat present in the epithelium as in a neutral 
condition. As to the fat in the intestinal cavity a great deal of attention 
has been given to determine whether it is present as a soap or an emulsion, 
but into a discussion of this question I do not propose to enter, and it 
suffices to say that there is abundant experimental evidence to show that 
both are present. In fact, to have a permanent emulsion, one must have 
present an alkali and some free fatty acid or, in other words, a soap, and 
as Heidenhain has pointed out, it is a common occurrence to see fat drop- 
lets of various sizes lying in the lumen. Indeed, it was the presence of 
such minute droplets in immediate contact with the striated border, 
together with the view I had taken regarding the structure of the striae 
as enunciated in the first part of this paper, that led me to think the fat 
was taken up in some particulate form and not alone in solution. 
In this connection, too, it is of interest to note that von Thanhofer* 
described a to and fro movement of the striae in the duodenum of winter 
frogs whose nerve roots had been severed. He claimed to have seen red 
* Pfliiger’s Arch., Vol. 8, p. 406. 
