1908-9. THE STELE OF OSMUNDA CINNAMOMEA. 519 
B.—Just above the junction of the first leaf-trace there are four 
tracheids in the cauline stele and three in the foliar. This number in- 
creases, but is very fluctuating, at the fifth node there being only nine in 
the one and three in the other. The first four leaf-traces answer to the 
¥Y% arrangement, the later ones to the 3g. The first root arises near the 
fourth node and the xylem of the stele remains wholly undisturbed in 
shape up to the seventh node except for a slight flattening at the sixth. 
C.—The phyllotaxy of the first six leaf-traces is approximately sym- 
bolized by % and the roots which are in a plane at right angles to that 
passing through the leaf bases are similarly arranged. ‘The 34 arrange- 
ment begins with the seventh leaf. The first disturbance in the stele 
occurs at the seventh node, where there is a flattening above the node. 
An eccentric parenchyma pocket of three or four cells is formed just 
below the ninth node, but the ninth leaf-trace is cut off by a band of 
parenchyma as in the tenth, and in neither case is there a gap nor any 
indication of a pith. 
D.—tThe arrangement of the first five leaves is of the 14 type; with 
the sixth begins the 3g. The first root comes off at the third node, and 
the second below the sixth. There is no change in the shape of the stele 
up to the eighth node. 
F.—The first seven leaves are arranged in two lines on opposite 
sides of the stem, the 3g arrangement begins with the eighth leaf. The 
roots occur without a break up as far as the apex of the stem, that is 
through seventeen nodes, a root for each leaf, and in a plane about at 
right angles to the radius of the stem that cuts through the corresponding 
leaf-trace. The xylem of the cauline stele at the first node consists of 
thirteen or fourteen tracheids, the leaf-trace of six or seven; at the third 
node the former consists of about twenty-five tracheids and the latter of 
ten. There is a flattening of the xylem of the stem at the third, fourth, 
and fifth nodes; at the sixth node there is a laterally-placed pocket of 
two or three parenchyma cells, that opens out as a bay or depression in the 
xylem, just above the connection of the leaf-trace. At the seventh node 
there is again a pocket of three or four cells, and a corresponding V-shaped 
depression just above the point of origin of the foliar strand, and into 
which the phloem tends to dip. The xylem soon rounds up again, but at 
the eighth node there is a repetition of the phenomena described for the last 
two, except that when the gap closes up, a few parenchyma cells are 
included. A narrow pith of four or five cells now persists up through the 
tenth node, with a gap corresponding to each leaf-trace. ‘The phloem is 
always most abundant opposite these gaps. At the eleventh node there is 
