6 STUDIES IN ROOT-PARASITISM. 
median strand is continued almost to the surface of the host’s 
root. It would appear that this is the first stage in the devel- 
opment of the gland. 
5. In the next stage figured, the gland is already well 
formed (Plate II, fig. 2). The arrangement of the outer, clear 
and inner, starch-laden cells of the cortex closely resembles 
that in Olax, but the cell contents have not been drawn in the 
figure. At the line of separation between these two parts ot 
the cortex, however, the collapsed layers are becoming appa- 
rent (bb). And here we meet with another marked disagree- 
ment with the course of development in Olax. The collapsed 
layers in Cansjera are developed like those in Santalum. 
The curved lines of growth are well seen. The nucleus is 
sharply separated off from the tissue outside it, and the vas- 
cular loop is indicated. The portion of ground tissue above 
the nucleus and between the arms of the vascular loop has cells 
yet more elongated horizontally than in the preceding stage, 
and there is evidence of pressure from the expanding nucleus 
below. 
In the lower part of the nucleus the gland is well seen. 
In the few examples obtained this organ is extremely simple 
in structure, consisting of a median split in the nuclear tissue, 
lined by secreting cells. The gland in Cansjera is totally differ- 
ent from that in Olar. It resembles that in Santalum in being 
lined by a well-marked epithelium, but it is doubtful whether 
a duct is present, as in that case. The lumen is narrow and 
uniform in all the cases met with. The glandular cells them- 
selves are like those already described for Santalum (Sant. I, 
para. 14). They have a dense mass of granular protoplasm 
at the base, while the upper part of the cell is clear (fig. a). 
The secretion, formed in this upper part of the cell, exudes 
in granular mucilaginous rods which fill the cavity of the 
lumen. These rods are better shown in the section drawn on 
Plate III (fig. 2). In the horizontal section through a slightly 
older haustorium (fig. 1) the various tissues referred to above 
