26 EXPERIMENTS IN THE HYBRIDISING OF INDIAN COTTONS. 
coloration.* In Stocks (Mathiola) yellow is recessive, as Correns 
also found in crosses between a white Polemonium ceruleum and 
P. Flavum, and in these the colour is contained in plastids. + 
In Verbascum blattaria L., Shullt finds yellow dominant, and 
here the colour is dissolved in the cell sap. Microscopic examina- 
tion shows that with cottons in which, too, yellow is dominant it 
is a sap colour. 
The neglectum (arboreum) shape of leaf is dominant over the 
herbaceum, and if we suppose that here again it is the phylogen- 
etically older character which is dominant, this fact would confirm 
einal 
Sir George Watt’s view that “ G. avborewm Linn. if not an ori 
wild stock, and therefore botanically a species, 1s remarkably near 
to what we are justified in believing, may have been one of the 
ancestral stocks of many of the cultivated cotton plants of the 
Old World § and that by cultivation was produced from the origi- 
nal G. arbsoreum, the annual plant now known as Gt. herbaceum. ”| 
The seed-coat does not appear to behave in exactly the same 
way : looking at a collection of seeds, from each plant of a bed, 
one would say at once that the ‘fuzzy ” and ‘‘naked” characters 
obviously segregate. But closer observation shows that there 
are intermediate grades of ‘“ fuzzyness.” It may be that these 
characters really do segregate in the same way as other varietal 
differences, but are influenced by external conditions, for 
Fletcher* found that in cottons of other species irrigation tends 
to make the seeds naked.** Or it may be that these characters 
ave “ poikilodynamic,” and while separating in the germ cells, as 
* Bateson (W.). Saunders (Miss E, KR.) and Punnett (R. C.) Reports to the Evolution 
Committee of the Royal Society II and III, 1905 and 1906. 
+ Correns(C.). Weitere Beitrage zur Kenntniss der dominerenden Merkmaale und der 
Mosaik-bildung der ‘‘ Bastarde ’ Ber. Dentsch. Bot. Gesells, 21: s.195—201. 1903, quoted 
by Shull (13). 
+ Shuli (13), p. 115, 
§ Watt (1), p. 86. 
| Watt ibid, p. 323. 
{ Fletcher Review of Sir George Watt's Book (1) Nature, Jan, 1908: Vol. 77, p. 242, 
** Among Hgyptian cottons Balls find2,on the other hand, much fuzz dominant over little 
fuzz, and Fletcher (5) in another pair of Indian varieties that fuzz is dominant Neither worker 
vives details of more than two generations. 
