144 THE MENDEL JOURNAL 



grossulariata) is crossed with its lighter coloured 

 variety lacticolor. Before the series of experiments 

 which we are about to describe were made by Mr. 

 Doncaster, lacticolor was known to occur only in 

 the female form, for males were unrecorded. When 

 lacticolor female is crossed with grossulariata male, 

 the offspring consist of the grossulariata form alone, 

 there being males and females in approximately 

 equal numbers. It is clear from this result that 

 lacticolor is recessive, since it does not appear in 

 the F^ generation. With regard to the dominance 

 of one sex or the other, we may for simplicity 

 regard the female as being dominant and as carry- 

 ing maleness recessive, as we did for Bryonia (supra 

 p. 134). The appearance of both males and females 

 in the F^ generation is then quite consistent with 

 that assumption. Let us now see how the other 

 possible crosses fall into line with such a scheme. 



If this representation is right, then the F^ 

 grossulariata males will be aU G L ^ ^^ and all 

 the females will be G L ^ '^ . The grossulariata 

 character being dominant, both these forms are 

 visibly grossulariata, but the male character being 

 recessive can only manifest itself when femaleness 

 is absent ; hence the first form is a pure male and 

 the second is visibly female, but carries male recessive. 

 It is necessary before we go farther to make another 

 working assumption. It is, however, not only ade- 

 quately supported by the facts of these particular 



* Where G — grossulariata, L=lacticolor, <S = male and $ = female. 



