200 THE MENDEL JOURNAL 



mated to the pink-eyed male. But if the reciprocal cross be made, 

 namely, " pink female " by " vermilion male," all the first genera- 

 tion, both males and females, are red-eyed. 



If vermilion-eyed females be mated to pink-eyed males, the F^ 

 generation has all its females red-eyed and all its males vermilion. 

 The second hybrid (F2) generation bred inter se from F^^ consists 

 of red-eyed males and females, vermilion-eyed males and 

 females, pink-eyed males and females, and orange-eyed males 

 and females. But when in the second or reciprocal cross, the 

 vermilion-eyed males are mated to their pink-eyed sisters, the 

 Fj^ generation are all red-eyed. The Fg generation derived 

 from F^ crossed inter se, consists of red-eyed and pink-eyed 

 males and females, and of vermilion-eyed and orange-eyed 

 males. That is, the females do not manifest either the vermilion 

 and orange eye colour ; these are the two eye colours which 

 are devoid of the pink factor. The females in this experiment 

 are therefore never devoid of the pink factor. Thus in the 

 F2 generation of this reciprocal cross we see again the 

 intimate relationship of femaleness and pinkness. Some of the 

 males are without pinkness, the females not. 



The same female sex relationship with pinkness is shown in 

 another way. In certain kinds of crosses, the offspring give only 

 half as many pink males as females, and twice as many orange 

 males as females. 



Professor Morgan endeavours to explain the association of 

 femaleness with pinkness, by assuming that the factor which 

 determines the manifestation of the pink eye colour, is carried 

 only in the same chromosome that also carries the factor 

 which determines the female sex. 



The rorm of Nose and its Segregative Inheritance. 



The form of a nose doubtless depends upon many factors. But 

 chief among them we may suppose are the length, breadth, and 

 angle of inclination of the nasal bones ; the form, length, breadth, 

 and thickness of the nasal septum, and the degree of development 

 of the turbinal bones. The segregation and persistence in 

 families of a definite type of nose-form is a subject well worth 

 further study. The inheritance of this character from the 

 Mendelian standpoint has not yet been adequately studied. 



We are able in the accompanying photograph to show what 

 appears to be an undoubted transmission of a very prominent 

 form of nose from a grandmother to a grandson. The grand- 

 mother (on the right of the photograph, who is now over eighty 

 years of age) was the wife of a gipsy and she herself came of gipsy 

 stock. She and her husband eventually settled in a small village 



