HOWARD AND HOWARD. 1 5 



A marked difference between the short hairs in Pusa 20 and 

 those in Punjab Type 9 was noticed. In this cross in the wheats of 

 the gametic constitution S s 1 1 the short hairs almost disappear 

 and it is only with the greatest difficulty that these forms can be 

 distinguished from those with smooth glumes. This extreme 

 reduction of short hairs in the Fi was met with again in one of the 

 cases of simple felting referred to below. It would therefore appear 

 probable that the short hairs in Pusa 20 are not due to the same 

 factor as those in Punjab Type 9. It is also improbable, judging 

 by appearance, that the long hairs in both these wheats are identical, 

 the long hairs in Punjab Type 9 being much denser and longer 

 than those in Pusa 20. It is therefore probable that we have four 

 different kinds of hairs, each due to an independent factor and 

 inherited separately, but this can only be proved by further experi- 

 ments. In Pusa 20 both the kinds of hairs do not differ very widely 

 and it will be much more difficult to analyse the F 2 generation 

 than in the crosses with Punjab Type 9. 



Simple Felting. 



In two of the parents with felted chaff only one class of hairs 

 was distinguished under the microscope, and in both eases the 3 : 1 

 ratio was obtained in the F 2 . The felted parents in these crosses 

 were lines selected from common wheats and are described in the 

 records as B XI 77 and Pusa 4. The hairs on these wheats, when 

 directly examined, appeared to be different, and this was confirmed 

 by their behaviour in the Y x . In the Fj from Pusa 4 the same pheno- 

 menon as in the crosses on Pusa 20 was observed, namely, the almost 

 total disappearance of the hairs, while in the F, from B XI 77 the 

 felting could be seen by the naked eye. If the felting factors in 

 these two wheats are different, a cross between them would give 

 smooth progeny in the F.. The actual experiment, however, has 

 not yet been carried out. The details of the second generation of 

 the above crosses are given in the following table : — 



