128 STUDIES IN INDIAN COTTONS 



is given in Table 1\'. From this table it is seen that, of 15 eyed 

 parents grown under conditions admitting of cross-fertihzation 

 taking place natui-ally, 14 have given eyed forms only and are 

 therefore pure. One only has given eyeless plants in mimbers 

 which closely approximate to expectation on the supposition tliat 

 the parent is impure and presence of the eye is fully dominant over 

 absence. Two only of the eyeless parents have bred pure. Of the 

 •200 offspring of the remaining 4 plants 19 have the eye fully devel- 

 oped, while in the remainder it is absent, a number readily accounted 

 for by the amount of cross-fertiUzation knowii to occur, but incom- 

 patible with any simple Mendelian ratio. In a single case only 

 has a form appeared in which the intensity of the eye is definitely 

 reduced. This form is being submitted to furthei' investigation.* 

 It would appear, therefore, that the intermediate form of eye found 

 bv Balls does not occur in the present group.* 



(C). — The Red (■(>l()ioiH(/-)Ka(frr in the Sap. 



Type 3 and Type 11 are characterised l)y the picscnce of a 

 red anthocyanic colouring-matter in the sap which impaits to the 

 entire plant, stem, foliage and flower, a deep red coloui'. \\\ these 

 types the intensity of the red in the petal is sufficient to mask the 

 true petal colour entirely, and it is only in a few cases of diseased 

 flowers and of such as open out of season that the presence of 

 the vellow petal colour in Type 3 can be determined by direct 

 observation. When a plant, which breeds pure to this character, 

 is crossed by one in which the i(m1 coloui' is absent, the K, 

 generation bears the red colour which uiav be snid to be doiiii- 

 nant. 



The intensity of the i-ed colour is. howcxcr. definitely diiui- 

 nished and the petal attains a condition which has been denoted 

 in the case of a cross with a yellow flowered plant as " red on yellow.' 

 It is conse(|uently ])ossible to say, from an examination of the petals 

 of plants derived from such a cross, whethei- \\\v plant is pure with 

 regard to this character oi- not. H'roni an examination of Table V it 



* Since the above was -aTitten this form has been found to breed true to the intermediate 

 condition. It must, therefore, represent a further type. 



