BUTLER AND HAFIZ [97 



type ; in others again the narrowly filiform second type predomi- 

 nated, while there were numerous "intermediate" spores and 

 only a few typical bicellular brown spores ; finally in some cases, 

 especially in the earlier cultures, all the types above described, 

 except the " intermediate," were found, though 3-celled spores 

 were never common. 



Hence we have had some difficulty in deciding whether the 

 fungus should be considered to belong to the sections Phceodidymce 

 or Ph(eo2^hra<imice of the Sphcprioidacefe. But the characters as a 

 whole agree better with those of the latter section, of which 

 Hendersonia is the main type, than with the former, which 

 consists mainly of Dvplodia and its allies, and the S-celled brown 

 spores should, we think, be considered as the most highly difi:*er- 

 entiated and, therefore, most important for systematic purposes. 

 The genus differs from the rest of the Phwophragmiw chiefly 

 in the characters of the stroma and the possession of two distinct 

 types of spore in the same loculus. The latter peculiarity 

 approaches it to the genus Phomopsis amongst the Ilyalosjwrrr , 

 and it is of great interest as supporting the view that the 

 hyaline, filamentous bodies of the latter genus are true spores, 

 and not merely disjuncted basidia as has been maintained by 

 some observers. There cannot be the smallest doubt that these 

 bodies in the present fungus are spores, for they are borne on 

 true sporophores, in an exactly similar fashion to the spores of 

 the first tj^pe. Another fungus in which two spore forms are 

 found in the same pycnidium is Fusicoccum viticolum Red*. ; 

 which Shearf has shown to be the imperfect form of Cruptos- 

 pforella viticola Shear. Shear distinguishes the two spore forms 

 as pycnospores and scolecospores, and believes the latter to be 

 true spores, not basidia as held by Reddick. The pycnidia 

 themselves are immersed in a stroma, which is evidently of much 

 the same type as that of the sugarcane fungus.| It should be 



♦ Reddick, D. Necrosis ot the grape vine, Cnrnoil Univ. Agr. Expt. Stat. Bull. 26.3, 190!<. 



t Shear, C. L, The ascogenous form of the fungus causing dead-arm of the grape. Phyto- 

 pathology, I, 1911, p. 110. 



t Gregory, C, J. A rot of grapes caused by Crifptosporella ritirola. Phytopathology, 

 111,1913, p. 20, fig. 1. 



