~-MAGANLAL L. PATEL }25 
If, however, the accessory bud had developed first into a vegetative branch 
however small, which had then given rise to a sympodium, the estivation of 
the flower on such a sympodium would be in the same direction as on the’ 
primary fruiting branch arising from the main bud. In all the cases which 
we have examined, and such cases are very common in the upper part of the 
plant at Surat, the estivation of the flowers is found to be in the opposite direc- 
tion on the two primary fruiting branches arising from the node in one axil. 
This proves, we think, that in Gossypium herbaceum at Surat at any rate, both 
buds in an axil may form sympodia. Illustrations of such occurrences will be 
seen in Plates VII and VIII. 
The following example will make the point clear :— 
Suppose that the position of the accessory bud is on the right side of the 
main bud. The zstivation of the flower at the first node of the fruiting branch 
developing from the main bud, will be against the hands of a watch, while the 
stivation of the flower at the first node of the fruiting branch developing 
from this accessory bud will be in the same direction as the hands of a 
watch. 
In the case of the flower at the second node of the fruiting branch 
developing from the main bud it will be in the same direction as the hands 
of a watch, while in the case of the flower at the second node of the fruiting 
branch developing from the accessory bud, the estivation will be against 
the hands of a watch. This rule is applicable at all further nodes. 
The opposite is the case in both sorts of branches when the position of the 
accessory bud is on the left of the main bud. 
2. The leaf-factor. In connection with the various species of Gossy- 
pium, the term “ leaf-factor”’ was introduced by Leake! in order to define by 
a single figure the relationship between the breadth of the middle lobe of the 
leaf and the indentation of the leaf, and though some doubt has recently been 
raised by Kottur as to its simple character, yet it is a convenient figure for 
the purpose in question. There is a good deal of interest attached to the 
consideration as to how the shape of the leaf as thus described varies on different 
parts of the plant, and Leake has stated that in his observations the leaf- 
factor of the leaves on the primary fruiting branches is the smallest, on the 
main stem somewhat greater, while on the monopodia it is the largest of all. 
This relationship applies, according to Leake, to all types of cotton which he 
has examined, | 
So far as various strains of Broach deshi and Goghari types of Gossypium 
herbareum grown at Surat and Broach are concerned, we have found, however, 
1 Journai of Genetics, 1, p. 220, 1910-11, 
