186 THE INHERITANCE OF CHARACTERS IN RICE, II 
Four factors have been identified with certainty and one more, &, is put 
down provisionally. Adopting the ‘ presence and absence ’ notation, their 
description is as follows :— 
G produces dark gold. 
[ modifies all golden colowing to a corresponding degree of dark 
furrows and inhibits golden colouring of the internode. 
P produces a piebald pattern on dark gold or dark furrows. 
L gives tipped gold from dark gold, ripening straw from ripening gold 
and granular dark furrows of degrees. 
E regarded, provisionally, as giving even-colouring by prevention of 
mottling. 
In the absence of G@, and apart from other complications, the inner glumes 
are ripening gold. This colouration is quite distinctive, the glumes being 
almost green at flowering and passing through lemon yellow to a dull light 
gold at maturity (Plates I and II, fig. 1), unset grains remaining green. No 
case of the absence of this colouring, other than through inhibition, has been 
recorded, hence its production is not assigned to any factor in the notation 
employed. 
A golden yellow internode is present in all plants with golden glumes, as also 
in the ripening straw type where ripening gold is inhibited by 7. In other 
words, it occurs in all plants in the absence of J. 
The factor G. 
In the absence of J this factor produces a dark gold colouring of 
the inner glumes. The latter are definitely gold at flowering and darken 
as the grain develops to a reddish brown at maturity (Plate I, fig. 2, and 
Plate II, fig. 4). In unset grains the glumes do not darken appreciably 
with age. 
In the presence of J all trace of gold disappears and it is replaced by a 
blackish brown pigment, showing mainly in the furrows of the glumes, giving 
the type described as dark furrows (Plates I and II, fig. 5). From the results 
of a large number of crosses it appears that dark furrows and dark gold are 
produced by the same factor, G@, and that the presence or absence of J deter- 
mines which colour shall develop. 
A similar modification takes place in the case of ripening gold which, in 
the presence of I, is replaced by a very dilute form of dark furrows described 
as ripening furrows (Plate I, fig. 4, and Plate II, fig. 2). Thus G may be 
simply an intensifying factor which darkens the ripening gold and furrows 
types. 
