196 THE INHERITANCE OF CHARACTERS IN RICE, I 
varieties are fine. The distinction is more a matter of the relation between 
length and breadth of the grain than of absolute measurements, though very 
commonly GG varieties are both shorter and broader than gg varieties. 
Crosses between the two types give an intermediate F, and an F, com- 
prising three groups—coarse @ G : intermediate Gq: finegg imal: 2: ratio. 
The connection was first noted definitely in a family, No. 1247 N, giving 
a 3:1 ratio of dark and medium gold: ripening gold. Grain measurements 
were made for each plant and the averages for the two groups were as follows :— 
No. of Length Breadth Length 
plants mm. mm. Breadth 
Dark and medium gold ss 1182 8°45 3°00 2°8 
Ripening gold af oe 378 8°85 2p 3:2 
In all cases, both here and in later figures, the grain measured was the 
ordinary grain of paddy that breaks off on threshing. This comprises the rice 
grain enclosed in the inner glumes, with the receptacle and outer glumes 
attached at the base (Plate III, figs. 1-6). It was found that the measure- 
ment of three of the end grains from a well-developed panicle was sufficiently 
accurate for each plant and, except where otherwise stated, the figures that 
follow are based on such measurements. 
Further families from the above lot confirmed these results : pure dark 
golds were coarse, pure ripening golds were fine, and splitting lots gave three 
groups— coarse dark gold, intermediate medium gold and fine ripening gold. 
The averages of several pure lots are shown in Table VIII compared with 
one of the splitting lots. The average of a number of natural crosses, occurring 
as medium golds in the pure ripening gold lots, is also shown. 
Tasie VIII. 
Length and breadth of grain and factor G. 
a 
Dark gold Medium gold Ripening gold 
Ref. No. Character o 
parent = 
L. | B. L. B. L. B. 
| —— —- —_—____ 
1794 N Dark gold Sion 3°0 “ A 
1795 N do. 8°3 2-9 ah re 
1796 N do. 78 2°9 S ye E 
1789 N | Ripening gold te a a 8°8 2°6 
1790 N do. re as nes 8-9 2:6 
1791 N do. : 5¢ ne 9-0 2:7 
1792 N do. ae “fs Pe: i 8°9 2-7 
1800 N | Medium gold 8-4 3°0 es a3 9°] Me | 
ca Sie 8: 2° % 
Crosses (36 plants) 
N.B. Measurements are in millimetres. 
