PARNELL, AYYANGAR, RAMIAH, AND AYYANGAR 203 
AAnnyr. The latter was selected from a family that had given unpigment- 
ed plants with red rice, thns showing that the pigmentation factor lacking 
must be N. F, was pigmented, red, as expected from the constitution 
AaNnRvr. In F, five groups appeared as shown below. The figures 
are not as near as could be desired to expectation but there is no doubt 
that they represent the ratio shown. 
PIGMENTED UNPIGMENTED 
Red White | 2ed | Grey-brown White 
| 
| ANR ACN? 2 eA Bel 7 NON, An 
| | a 1 ) v aN jr 
| | an 
| : | 
= NS | | | 
No. 2508 at 2837) 119° | 108 | 129 85 
EE AON AS) SI BEN OAT ree 305 : 102 : 102 : 136 ° 70 
These results definitely confirm the above theory regarding the nature 
of grey-brown rice and also afford additional confirmation of the existence of 
two factors necessary for the production of anthocyan pigmentation. 
Red rice and golden rice. 
An account has already been given above of the factor I which inhibits 
gold in the internode and changes any form of gold in the glumes to a corre- 
sponding form of dark furrows. It appears that this factor also is necessary 
for the production of red rice by the factor R. In the absence of J, that is to 
say, in all types with golden glumes or internode, the factor R produces golden 
rice. All red-riced plants so far seen have shown some form of dark furrows 
colouring in the glumes. The same applies to the various shades of light 
reddish rice that are common. This colouring is only produced in dark 
furrows types ; in golden types it is replaced by a yellowish or cream colour. 
At the time of ripening golden rice is very easily distinguishable from red, 
as shown in Plate V, figs. land 2. After keeping for some time much of the 
yellow colour disappears and the rice becomes duller and more brown. 
At this stage, if seen casually, it might be mistaken for red though the two 
are still separable with certainty. 
Golden rice has occurred in a large number of families showing segregation 
for [ in the presence of R. In all cases the red rice of the dark furrows group 
has been replaced by golden rice in the gold-glumed group. An example cf 
this is seen in the results from a cross between dark furrows, red rice and 
