94 



INHERITANCE OF ANTHOCYAN PIGMENT IN PADDY VARIETIES 



We have not found this to be the case in all our types, many having somo 

 shade of red in the leaf-sheath, but a blue-black colour in tlic stigma, and 

 the evidence quoted below proves that in some of these cases this purple 

 stigma colour appears to be due to the interaction of three factors, and further- 

 more that the simultaneous presence of all three appears necessary for the 

 production of colour at all. 



The cases discussed here with reference to the stigma colour are the 

 same as those quoted above (Table I) in which the leaf -sheaths gave the 9 : 7 

 ratio. The parents, selected in 1912 and 1913, had reddish coloured leaf- 

 sheaths, red apicuius and blue-black stigmas, and in 1913 and 1914 gave 

 the results below : — 



Table HI. 



If we again adopt the usual notation for such cases, the above results 

 could be produced by supposing that (1) the colour in the Icaf-sheath and 

 apicuius is due to a colour factor R acting on a Chromogen base C, (2) that 

 the purple colour of the stigma is due to a further factor P not piescnt in the 

 leaf-sheath and apicuius, and (3) that the sinmltaneous presence of all three 

 factors RCP is necessary for the jDroduction of colour of any sort in the 

 stigma. 



