KDiTii i;. s\i'Mii:i;s. tiik i;i;i:i;iiim. m- doi i;i.i: ri.nwKiis. .-,!i;) 



iiiilij II lilllc iiiorr lltiin liiilf llir F, crossljrcils give a mixitircor siiii^lcs and 

 (IdiiMcs, Hic n'iiiniiiili'i- lircod Inic hi sinti^lcnoss — licnro ni'iifli/ linl/' l/ir n/v/A'x 

 ill d-sinylcs luiisl m rnj ximjlrni'^s: l/ir ri'fil riirrij i/uiihli'urK^. 



The pr()|iiprli()i\ of douilles (icciirrini; iii llie nii\("(l l'a mi lies IVoiii ri II ici- Uiml 

 of maling was round in iiiosi oC Ihc unions lo bc I in 'i ; in soinc combinah^jns 

 •however il was <l('ai- llial aUlinni^h somo familios sIkiwimI Ihc iisnal ratio of 

 T) s : I (1, ollicr sisicr l'aniilics çonlaincd a vcrv nmch siiiallcr |iroporlion ol' 

 douilles. 



Wo are no\v in a position lo show liow Ihcso varions resuJts can be broughl 

 inlo Une and explained. NN'c arc cxidcnlly dealini^' hore willi a rase in whi(di 

 Ihe l'actors couccruiMl ai-c rcl.dcd lo ono anolhcr in a pccnliar lud dcfinitc nian- 

 nor, the nainrc ol' Ihc r(dalioii dcpending u|ion whclhcr llicsc l'arlors arc inlro- 

 duced inlo the individnal associaled togelher in onc ol' Ihc Iwo germ cells giving 

 rise lo Ihal individnal, or whelher they hâve been rcccixcd nome from Ihe onc 

 fonlribnting gei'm anA ^nme froin Ihc iil/icr. Il is riirllicr évident IhaL a sr.r- 

 liinilpil distribution ol' thèse facloi's among Ihc giTni cclls occiu-s in certain cases. 

 As regards this kind ol' interrelalionship between certain l'actors the Stocks 

 afVord a case parallel ^^ilh that reccntly described by Bateson and Punnetl in the 

 Swcet Pea {ljil/iijrii><) ' ])ul the liniilalion ol' Ihc powcrof cari'ving thèse faclors 

 to the gameles of onc scx iniroduccsan a<ldilional coinplcxilN in Ihc présent 

 case. 



W'e niay slatc the scvci'al conclusions arrived al in Ihc présent case as 

 l'ollows-. 



1) Singlencss is due lo Ihc |ii'i>seiiee ol' Iwo l'ai'loi's ( .\ and \). 



"î) Doubleness is due lo Ihe absence ol' cither oi' both ol' Ihcse t'aclors. 



.") in Ihe non-double-tiirowing single type X and Y are linlied together — 

 represenled IhusXY — so thaï when snch singles are i'ertiliscd vith llie .r// 

 gameles of the eversporting single, Ihegamelic recombinafions X (/ and .'Y can- 

 not be formed''. A gametic séries in F, plants thus derived, composed ol' cipial 

 numbers of XY'and xy gamètes, will llius give Ihe observed ratio of ." .s : ! rfin Fj. 



i) In the eversporting single the faclors X and Y are nol Ihns linked, but 

 the relation between Ihem is of the nature described ahove. Ihe parlicular com- 

 binalions of allelomorphs from which the eversporting zygote was ilsell' derived, 

 viz. XY and ./■// being those which occur most frequently among ils gameles, 

 vvhereas the recombinations X ,'/ and ./Y are fourni iiundi more rarelv. 



"i) .Vil four combinations of the allelomorphs riz. \\ . Xi/. .r\, .ri/ occur 

 among Ihe o\ules in an eversporting single, bul Ihc |iollen appears unable lo 

 carry X and Y eilher alone, or togelher. W'e hâve in facl hère a case of whal 

 bas been termed partial coupling 'combined witb a sex-limited distribution of 

 the faclors. 



The gametic séries in Ihe Iwo l\pcs of singles may bc rcprcsenlccl Unis. 



I. On Ihc iiilin--ivliilioMS (if lienrlic F.icldi's. Pr.ir. Uni/. Snr. II. \„\ II. l'.ljl. 



i. Il Wdiild hc iiiipossllile williin llic limils (illiic |ircsciil pipcp l(i ^iic Ihc cviin ii I;il diiln 



iipoii which thcsc cniiclusions are hased. A l'ull slalemeiil oC Iho resulls iihlaincd ami ;i ili^i-ii>si(iM o\ 

 llip samo are iiow in Ihe pi-ess, aiid mider llie title ■• Fni'lhci- Expcrinionis on Ihc Inhii I|;mhc (iT jiunlil- 

 inn and olher Cliaraclois iii Storks ', will apprar in Ihe l'orlhcnniing: niiinlier ni Ihc Jiuinuil „f Cm,-- 

 ticx (vol. i, 11° 4). 



7>. In a certain propcrlinn ol' i:i/iiirli;l s'iifïh's. i.i', lo saj sin^ilcs wliich ari' nol ol' piirc-luvd 



ofifçin linl derived l'roni inaliiit;s wilh cvcrspoi-liiip forins, \\c iiinv Mip|i(isc ih.-il Ihc jv^nic, il oh 



bi-ecdin<; liue to sinplcness, \\ill hum sonic {;.iincles m «hicli \ .mil V aie nul linhcd l(i;;cili.'r. 



4. See Datoson and Pnnn.'ll, Inr. ni. 



