k. MARTIN LEAKE Xkl) RAM PERSHAD. 'JO 



'these forms probablj' constitute a link — lacking in the Indian 

 cottons — between the monopodial types, in which the secondary 

 branches are pure monopodia and the sympodial types with pure 

 sympodial secondary branches. From the description given by 

 Todaro of his G. herbaceum it is clear that the plant to which 

 he refers is a sympodial type. He says they (the branches) 

 "grow in a zig-zag tortuous manner, one internode making a 

 small angle with another," a description which accurately defines 

 the form of a sympodial branch, but from his description of 

 G. wiyhtiamim it is not clear whether this plant has monopodial 

 or sjMnpodial secondary branches. There appears to be small 

 doubt that the Persian types approach, and in certain cases very 

 closely approach, the form grown in Southern Europe and de- 

 scribed by Todaro as G. herbaceum. The form from S. Europe 

 is only known to the authors from an entire plant preserved in the 

 herbarium of the Botany School, Cambridge University ; in this 

 only sympodial secondary branches are developed. 



In this diversity of the form of secondary branching is to 

 be found the most distinctive character of the various Persian 

 typeSj but in one other feature also they appear to differ with 

 some definitene.s.s. In all observed species of Gossypia there 

 occur two buds in the axil of each leaf, one of which is the main 

 axillary bud, and to this the other is accessory and lateral. This 

 accessory bud may remain dormant or may develop and, in the 

 latter case, gives rise to a monopodium (Plate VI). In certain 

 types development of the accessory bud appears to be a definite 

 feature, and in many cases the branch thus produced is far more 

 vigorous than that developed from the main bud. This is 

 especially marked wliere the main axillary bud forms a sympodium, 

 which is, in such cases, frequently reduced and in extreme cases 

 withered. The true relation of the branches is, here, readily 

 determined by close inspection from the fact that the smaller 

 branch is sympodial ^a condition which has never been found 

 in the branch from an accessory bud — and from the relative 

 position of the two branches. As has been shown, the position 

 of the accessoiy bud in relation to the main axial bud is constant 



