ICONES FILICUM SINICARUM 
PLATE 186 
LEUCOSTEGIA IMMERSA (Wallich) Presl 
POLY PODIACEAE 
LEUCOSTEGIA IMMERSA (Wallich) Presl, Tent. Pterid. 95 t. 4, f. 11 (1836); Hk. Gen. Fil. t. 
52A (1840); J. Sm. Hist. Fil. 84 (1875); Bedd. Handb. Ferns Brit. Ind. 51 (1883); 
Cop. Phil. Journ. Sci. 34: 240, 252 (1927); C. Chr. Contr. U. S. Nat. Herb. 26: 293, 
331 (1931); Ind. Fil. Suppl, III. 120 (1934). 
Davallia immersa Wallich, List no. 256 (1828, nom. nud.); Hk. Sp. Fil. 1: 156 (1846); Fil. Exot. 
t. 79 (1858); Hk. et Bak. Syn. Fil. 91 (1865); Clarke, Trans. Linn. Soc. II. Bot. 1; 443 
(1880); Christ, Farnkr. d. Erde 302 (1897); C. Chr. Ind. Fil. 211 (1905). 
Acrophorus immersus Moore, Proc. Linn. Soc. 2: 286 (1854); Ind. Fil. 2 (1857); Bedd. Ferns S. Ind. 
t. rr (1863). 
Humata immersa Mett. Fil. Hort. Lips. 102 (1856); Diels in Engl. u. Prantl; Nat. Pflanzenfam. 1: 
4, 209 (1899). 
Rhizome thick, woody, wide-creeping, hypogezous (subterraneous); scales linear- 
lanceolate, thin, rusty brown, sparce or rather copious at growing tip or base of stipe; 
frond far apart, stipe 20-35 cm long, firm, erect, dark-straminous or pale colored, smooth, 
nitente, glabrous from base upwards, lamina deltoid-ovate, acuminate, 25-35 cm long, and 
nearly as broad at base, tripinnate, pinnae about 10-jugate, oblique, long-petiolate, 
alternate, the basal pair much the largest, elongate-deltoid, 12-20 cm long, 6-10 cm broad, 
bipinnate; pinnules about 10-jugate under pinnate acuminate apex, the posterior basal 
one much the largest and produced, all petiolulate; ultimate pinnules I-1.5 cm long, to 
I cm broad, subrhombic-ovate, sessile, cuneate, apex roundish, lobato-incised with 
2-3 ovate lobes with bluntly dentate teeth; the upper pinnae narrowly oblong-lanceolate 
and gradually shortened; texture herbaceous, pale green, glabrous in all parts; veims in 
ultimate pinnules fine, repeatedly branched, one to each tooth, but not reaches tip and 
ended in a clavate hydathode; sorz large, 1-2 to each ultimate pinnule, terminating veinlet 
near the margin; indusiwm large, semi-orbicular, gray, membranaceous, entire, persistent, 
free on all sides except the lower side. 
Yunnan: Shweli-Salween divide, G. Forrest 25329, Szemeo, Henry 10083A; Mengtze, 
Hancock 63 (Kew No.); Yunnan-sen, Ducloux 1326, 6529; between Tengyueh and Lungling, 
Rock 7240; east of Tengyueh, Rock 7618. 
Also North India generally, Indo-China, Siam, Formosa, Luzon, the Philipping 
Islands and Malesia-Polynesia. 
This distinct fern resembles none of the genus, to which it belongs, by subterranean 
habit, pale green leaves with broad segments and large membranaceous indusium. 
Plate 186. Fig. 1. Habit sketch (natural size). 2. Portion of frond, showing venation and 
position of sori(x 10). 3. Scale from rhizome (x 27). 
