Phylogeny of the Pierinee. 299 
Huterpe theano and EH. ewrytele present us with another 
kind of departure from the ordinary Pierine condition, 
the latter being an excellent copy of Lycorea atergatis 
and Ceratinia (Ithomia) dionea; while EH. theano ex- 
hibits the form of H. ewrytele with a coloration much like 
that of Pereute charops ¢. 
We find then that, thus far back in the growth of the 
Pierine stock, a large section has been diverted under 
the influence of mimicry from the regular course of 
development of the Pierine pattern; and in order to 
continue the history of the latter, we must retrace our 
steps and take up the thread where we left it before 
beginning to discuss the mimetic forms of Pereute and 
Huterpe. 
Although it does not appear that the Old World can 
show any truly Pierine form so ancient as Huchevra 
socialis, there does exist a genus which seems to repre- 
sent in the Hast nearly the same stage of development as 
Catasticta, Leodonta, Pereute, and Huterpe in the West. 
This is the genus Delias, the close relationship of which 
with the western forms mentioned has been indicated by 
Doubleday and Westwood (Genera of Diurnal Lepi- 
doptera, vol. i., p. 33), and more distinctly suggested by 
Wallace (Trans. Ent. Soc. Lond., 3rd series, iv., p. 344).* 
The pattern of Delias belladonna is essentially similar to 
that of the early Catastictas, consisting as it does of a 
dark ground-colour relieved by paler touches in the 
interspaces, between which touches the ground-colour is 
already beginning to show a rudimentary division into a 
marginal and submarginal chain with a vaguely indicated 
discoidal patch in each wing. On the underside we have 
the same pattern in a slightly more distinct form; while, 
as in Catasticta, the pale areas of the hindwing and the 
apical region of the forewing are furnished with yellow 
streaks. ‘The red basal patches, so characteristic of 
Papilios have been led to meet the Pierines by discarding, or at 
any rate by not adopting, the bright metallic blues and greens that 
ornament the other sex. This was no doubt a shorter and easier 
way to the formation of an “inedible association,” than would 
kave been the acquisition by the Pierines of colours more nearly 
resembling those of the male Papilios. 
* Delias agrees closely in neuration with Pereute and Leodonta. 
See Butler, Cist. Entom., i., p. 40. Staudinger and Schatz, “ Exot, 
Schmetterl.,” 1892, Theil. ii., p. 63. 
