( V ) 



before us it is impossible to doubt the significance of the 

 diiierences which separate the female of manlius from its 

 male. In the female of phorbanta the central blue patches 

 have entirely disappeared, while the blue submai-ginal spots 

 of the hind-wing have become increased in size and tr.ins- 

 formed into white, Fuithermore, the ancestral submaiginal 

 band of blue spots in the fore-wing has also been tiansformed 

 into white. These changes, with the exception of that last 

 mentioned, produce a rough mimetic likeness to Euplaea 

 youdoti, as may be seen in Colonel Manders' beautiful plate 

 (Trans. Ent. Soc. Lond., 1907, PI. xxix, figs. 6a and 1). It 

 is evident from Colonel Manders' account (1. c, p. 451) that 

 the resemblance which appears to be so slight in the cabinet is 

 much enhanced by the mode of flight, and the fact that Faj/ilio 

 and Eupl(ta inhabit the same localities. Nevertheless it is 

 impossible to be satisfied with the simple conclusion that the 

 female phorbanta has gained its present pattern under the sole 

 influence of Etiplcea goudoti as we now know it in Bourbon. 



The ancestral submarginal blue spots of the hind-wing of 

 the male pliorbanta are already somewhat larger than the white 

 spots occupying a similar position in Eiiphea goudoti. The 

 blue spots transformed to white cannot theiefore have under- 

 gone a further increase in size in the female under the 

 influence of the existing E2iphea. Nor is it possible to 

 account by the same influence for the submarginal white spots 

 of the fore-wing of jjhorbanta ; for Colonel Manders (1. c, p. 435) 

 only knows of a single specimen of goudoti " with faint but 

 decided indications of a submarginal row " in the fore-wing. 

 It is obvious that the present pattern of the Bourbon Euplcea 

 cannot affoid us the interpretation of the change which has 

 occurred in the female J'ajiilio. 



If the upper-surface pattern of the female p)horbanta, fig. 6a, 

 on Colonel Manders' plate, be compared with that of Salumis 

 aiigustina, fig. 3, and Euphra goudoti, fig. 1, it will at once be 

 seen that the Papilio bears a far closer resemblance to the 

 Nymphaline than to the Euplceine. Now the upper-surface of 

 the SaJainis is a fair mimic of the Mauritian Euphea eiiphone, as 

 was pointed out by Mr. Roland Trimen, F.E.S., in 1866: — "In 

 spite of the very different outline of wings, the general 



