and of several Brachyccrous families in Diptcra. 289 



labial, as I find atrophying blades of the niaxillaj at 

 their bases. In my paper on the mouth parts of the 

 " Nemocera," published in the " Journal of the Royal 

 Microscopical Society " in 1904, 1 described them as labial 

 as my prej^arations showed no maxillre, but my dissections 

 lately made have been more successful, and enable me to 

 say that the lacinia or blade of this insect is 85 i^. or 

 -^^jj inch long, and I submit that it is as easy to overlook, 

 as it is difficult to dissect, objects of this size. 



It will thus be seen that I have found a striking affinity 

 between these three families in the mouth parts, which 

 show that the true place of Pliora is among the older 

 families of the Brachycera. 



In the PhoridjE the palpi in the limited number of 

 species I have examined are labial. This point I very 

 carefully studied, as the palpi being maxillary in Lcptis 

 and Lonclioptcra made me suspect that I was wrong in 

 describing them as labial, though perhaps this has little 

 bearing on the matter, if it be remembered that Bihio and 

 Chironomvs differ in this respect. 



The palpi are very large, and bear a number of the 

 characteristic Phorid bristles ; at their bases are many 

 wrinkles, scarcely annulations, and in one preparation, 

 Phora concinna, Mg., they are two-jointed. This is not, 

 however, the only Nematocerous character; in T. atcrrima 

 the palpi bear the sense organ, found on the second joint 

 in the Mycetophilidte, Bibionida3, Simulidse and Rhyphidse. 

 This I have thought to be an olfactory organ and I have 

 figured it as such, as it is found on the palpi of Bihio 

 hortulanus, L.* 



9. The genitalia (male). The hypopygium is free and 

 is joined to the abdomen by a membrane ; it is furnished 

 with representatives of the usual paired organs. In P. 

 incrassata these are hairy bulbs much closer in function 

 and appearance to the cerci of Pcriplancta than to the 

 usual hooks or forceps of Diptera. In another species, 

 P. concinna, Mg., only a single hairy organ with a trans- 

 verse suture, placed on an asymmetrical platform, is present. 

 In T. atcrrima two minute hairy bulbs are present. The 

 part that supports these organs and the anus, is in a more 

 dorsal position and is separated by a suture from another 

 lower part. This lower part seems to be formed of the 



* Some new sense organs in Diptera. "Jonr. Qiiekctt I\[icr. 

 Chib," ser. 2, vol. ix, Plate VII, 1904. 



TRANS. ENT. SOC. LOND. 1908. — PART II. (SEPT.) 19 



