Rev. F. D. Morice's Notes on Australian Sawflies. 323 



gaeic forms than between the latter and any now to be 

 found in Arctogaea. 



As to " larval " characters, we have seen that occasion- 

 ally in Australia, but never in Arctogaea, forms which 

 feed moving freely about over their food-plants have 

 notwithstanding developed no pro-legs. It would be in- 

 teresting to know whether any such cases occur in Neogaea, 

 and especially if there are any among such genera as in other 

 ways seem to show affinity with Perga, etc. But I have 

 sought in vain to get any information on this point, so we 

 may pass on at once to consider the characters of imagines. 



(a) Venation of the fore-wing. 



i. In most groups of Arctogaeic Sawflies, and in almost 

 all those which may be considered typical and dominant 

 in that Realm, the radial cell is divided by a transverse 

 nerve. The exceptions are the Arginae, the Lophyrinae, 

 and a great majority of the Nematinae. 



On the contrary in Notogaeic forms, to whatever group 

 t hey may belong, the radial cell is invariably undivided. In 

 some cases this is not surprising, for three of the Australian 

 genera are Arginae, and others appear to be more akin 

 to that group and probably also to the Lophyrinae than to 

 any Arctogaeic genus in which the radial cell is divided. 

 But we cannot thus account for the absence of a transverse 

 nerve in the Syzygoniides (Perga, etc.). The only existing 

 Arctogaeic Family in which these could possibly be placed 

 is that of the Cimbicinae, and all Arctogaeic genera of that 

 Family have the radial cell divided. Such at least is now 

 the case though it is not easy to explain why it should be 

 so, for the earlier (fossil) genera of Cimbicinae — the Phena- 

 copcryiiii of Ivohwers Classification — are stated to have 

 the cell undivided, so that — contrary to what might have 

 been expected — the venation of modern Cimbex, Abia, 

 Amasis, etc. seems to be more "generalised" than that 

 of their probable ancestors. But it is possible, no doubt, 

 that the Syzygoniids and Phenacopergini represent one 

 branch of the Cimbicinae in which the "transverse radial 

 nerve " long ago disappeared, and the Arctogaeic Cimbicinae 

 another branch of the same stock which have retained it. 

 However, in any case, the universal absence of this nerve 

 in Notogaeic Tenthredinidae is a circumstance which 

 deserves to be noted. 



ii. A character which separates all Arctogaeic Sawflies 

 from Hymenoptera of other Orders is the presence of a 



