130 DR. E. KLEIN. 



Aegineta, Carmarina hastata, a peripheral " zone of granules^' 

 (Kornclienkreis), due to the above circular fibrils. In the case 

 of the nuclei of the epithelial cells of intestine and other organs 

 of mammals this arrangement is not of a general character, as it 

 is absent in many instances. I have seen it only rarely in the 

 nuclei of the ceils of stomach and mesentery of newt. But 

 whenever the nucleus contains next its membrane a regular row 

 of granules, it may be taken as certain that this is an index for 

 the circular tibrils. I am inclined to think the same of the row 

 of dots in the germinal vesicle of the ovum, represented by Bal- 

 four in fig. 21 on PI. XVIII, although Balfour does not notice 

 anything of a network in this germinal vesicle. 



As is well known, the epithelial cells covering the villi of 

 small intestine and the mucous membrane of large intestine, as 

 well as those lining the crypts of Lieberkiihn, possess at their 

 free border a fine longitudinal striation, which, as had been first 

 proved by Brettauer and Steinach,^ is due to its being composed 

 of minute rods. Thanhoffer" maintains for the duodenal epi- 

 thelial cells of frog that the above fine striation is the expression 

 of contractile processes of the protaplasm of the cells, which 

 processes are said to play an important part in the absorption of 

 fat. Portunatow^, however, while opposing this view of Than- 

 hoffer^s, mentions the presence of short protoplasmic processes as 

 cilia in the above epithelial cells of frog. 



In the case of the ciliated epithelial cells of the foregut of 

 newt, I have mentioned in my first paper that the cilia are dis- 

 tinct prolongations of the fibrils of the intracellular network, and 

 in this respect I was at one with Eberth, Marchi, Eimer, and Nuss- 

 baum. The same relation I notice also to exist with regard to the 

 striated border of the intestinal epithelial cells, viz. that the stria- 

 tion is due to Jine fibrils projecting a short distance beyond the 

 general intracellular netioorh (fig. 1) . I cannot definitely decide 

 whether the bright thick cuticle, which apparently lies on the 

 free surface of the epithelial cells, and which apparently contains 

 these fine strise, is really a cuticle covering the cells — as is repre- 

 sented by most histologists since that bright cuticle was first 

 observed by Henle — or is, on the contrary, only a substance 

 arranged aro2ind the free border of the cells without covering 

 the latter, i.e. is a projection of the interepithelial substance, as 

 maintained by Thanhoffer with Lenhossek. 



I know of one fact which seems to me to show in a clear 

 manner that the fine strige, i.e. the fine fibrils projecting from the 



* ' SitzuBgsber. d. Kais. Akadem. d. Wiss.,' 1S57. 

 2 Pfliiger's ' Archiv/ viii, p. 391, and ' Centralblatt f. med. Wiss.,' 1876, 

 No. as, p. 101. 

 '■" Pfliiger's 'Archiv,' Bd, xiv, p. 285, 



