( V ) 



before us it is impossible to doubt the significance of the 

 differences which separate the female of manlius from its 

 male. In the female of phorbania the central blue patches 

 have entirely disappeared, while the blue submarginal spots 

 of the hind-wing have become inci*eased in size and trans- 

 formed into white. Furthei'niore, the ancestral submaiginal 

 band of blue spots in the fore-wing has also been tiansformed 

 into white. These changes, with the exception of that last 

 mentioned, produce a rough mimetic likeness to Euplcea 

 goudoti, as may be seen in Colonel Manders' beautiful plate 

 (Trans. Ent. Soc, Lond., 1907, PI. xxix, figs. 6a and 1). It 

 is evident from Colonel Manders' account (1. c, p. 451) that 

 the resemblance which appears to be so slight in the cabinet is 

 much enhanced by the mode of flight, and the fact that Fapilio 

 and EujiloM inhabit the same localities. Nevertheless it is 

 impossible to be satisfied with the simple conclusion that the 

 female phorhcmta has gained its present pattern under the sole 

 influence of Eupdcea goudoti as we now know it in Bourbon. 



The ancestral submarginal blue spots of the hind-wing of 

 the male phorbanta are already somewhat lai'ger than the white 

 spots occupying a similar position in Etiphea gotidoti. The 

 blue spots transformed to white cannot thei-efore have under- 

 gone a further increase in size in the female under the 

 influence of tho existing Euphea. Nor is it possible to 

 account by the same influence for the submarginal white spots 

 of the fore-wing of phorbanta ; for Colonel Manders (1, c, p. 435) 

 only knows of a single specimen of goudoti " with faint but 

 decided indications of a submai'ginal row " in the fore-wing. 

 It is obvious that the present pattern of the Bourbon Euploia 

 cannot affoid us the interpretation of the change which has 

 occurred in the female Papilio. 



If the upper-surface pattern of the female p)horhanta, fig. 6a, 

 on Colonel JManders' plate, be compared with that of Salamis 

 augustina, fig. 3, and Euploe,a goudoti, fig. 1, it will at once be 

 seen that the Papilio bears a far closer resemblance to the 

 Nymphaline than to the Euplu?ine. Now the upper-sui'face of 

 the Salamis is a fair mimic of the Mauritian Eupleea euphone, as 

 was pointed out by Mr. Eoland Trimen, F.R.S., in 1866: — "In 

 spite of the very different outline of 'wings, the general 



