27 



From the above description it can be seen that, 

 although water may enter the mantle cavity all along the 

 external opening of the mantle, yet when the funnel is 

 locked the only way out for the contents of the mantle 

 cavity excretory or genital products, water, &c. is 

 through the central funnel chamber. As in oilier 

 Octopods, Eledone has no valve in the funnel. It has, 

 however, a large and elaborate mucous gland — Mhller's 

 gland (PI. II, fig. 10, f.o). This is four-lobed, and is an 

 elaboration of the internal epithelium of the funnel, and 

 may best be seen by opening the funnel ventrally, as in 

 PI. II, fig. 15. It serves to lubricate the internal surface 

 of the funnel, which consequently is generally coated over 

 with opaline viscous mucus, rendering the gland itself 

 rather obscure in fresh specimens. It may be seen, how- 

 ever, on scraping the mucus away. 



Pallial Complex (figs. S and 11).— Under this general 

 term may be included those important organs situated in 

 the mantle cavity, together with the external apertures 

 of certain internal organs. Eledone, like other Cephalo- 

 pods, in spite of its high specialisation along certain lines, 

 has yet retained its primitive symmetry in certain 

 features, including the pallial complex. The organs of 

 the pallial complex are : — 



(1) A pair of Gills, one on each side of the visceral 

 mass, and attached to it by muscles, vessels, &c. 

 (fig. 11, .'/•>; 



(2) The Anus, situated anteriorly, between the left 

 and right halves of the vertical septum (fig. 11, an.) ; 



(3) The Urinary papillae — one pair, protruding for 

 about 12 mm. in a large specimen into the mantle cavity, 

 just in the angle between the base of the gill and the 

 visceral mass (tig. 11, JJr. //.). The urinary aperture is a 

 small hob 1 at the tip of this papilla : 



