4 BULLETIN 132, UNITED STATES NATIONAL MUSEUM 



characters which would place it in the Olethreutinae did not genitalia 

 and the notched termen of fore win*? show its clear relationship to 

 the Epinotia group of the Eucosminae. In the Olethreutinae, Endo- 

 thenia, and Ema exhibit the venational characters of the Laspeyresiiae 

 (vein 5 or hind wing parallel to 4) . They also have the general hab- 

 itus of the latter ; but the genitalia are typically Olethreutin, Endo- 

 thenia having a strongly developed uncus (not found in Laspeyres- 

 iinae), and both genera, the heavy spine cluster on sacculus of harpe 

 typical of the Olethreutiinae. Endothenia forms (with Gijmnan- 

 drosma and Ecdytolopha in the Laspeyresiinae) the connecting link 

 between the two subfamilies. Episimus, which is here referred to the 

 Olethreutinae might go, on the other hand, as easily into the Eucos- 

 minae were it not for the normally connate condition of veins 3 and 

 4 of hind wing. Both the subfamilies Laspeyresiinae and Eucos- 

 minae seem to be derived directly from the Olethreutinae which ap- 

 pears to be the primitive group of the family. 



The trees shown on pages 7 and 77 illustrate my conception of 

 the relations and phylogeny of the genera of the two groups here 

 treated, and the tables opposite pages 6 and 76 give comparative 

 analyses of the various structural characters upon which the genera 

 are classified. It will be necessary here only to expand the original 

 key to the subfamilies ** by a brief summary of the characters dis- 

 tinguishing them. 



CHARACTERS OF THE SUBFAMILIES OF OLETHREUTIDAE 



Olethreutinae. — Hind wing normally with vein 5 bent at base and 

 approximate to 4, veins 3 and 4 connate. (Where 5 is straight and 

 parallel with 4, harpe of male genitalia has at least one strong spine 

 cluster {Spc^) always present; and where 3 and 4 are stalked 

 {Episimus tyrius) there are long flat spines on base of sacculus). 

 Thorax normally with strong posterior tuft (absent only in Episl- 

 Qiius ttTid Bactra). Fore wing with termen normally convex, rarely 

 concave {Episimus) and never notched; no costal fold in male. 

 Male genitalia with uncus normally present and well developed, 

 usually simple ; socii usually well developed ; harpe with at least one 

 and frequently two strong spine tufts {Spc^, Spc-) from or near 

 sacculus; cuculli usually narrowly elongate; sacculus often with 

 strong basal spining {ScSp). 



Eucosminae. — Hind wing with vein 5 always somewhat bent and 

 approximate to 4 at base; veins 3 and 4 stalked or united (connate 

 only in Gwendolina where termen of fore wing is notched, and sac- 

 culus of harpe is haired but without strong spine cluster). Thorax 

 seldom Avith posterior tuft. Fore wing Avitli termen convex, con- 



» Bull. U. S. Nat. Mus. 123, p. 10, 1923. 



