LEJEUNEA. 71 
perianths, &c., also closely correspond. The Oriental Br. fruticosa, Lg. et 
G., however, I have been obliged to separate as a distinct subgenus, for 
the 2 branches are innovant, sometimes forked, and the perianths 
obcordato-triquetrous, with cristate keels and one or two ridges added on 
to front and back. 
A Bryopteris of Nees (Br. spathulistipa), anda few new species, consti- 
tute the genus Thysananthus, Ldng. in ‘Syn. Hep.’ The character on 
which it is mainly founded—the fringed keels of the trigonous perianth— 
is neither very strongly marked nor very constant; but the plants have 
another and more important character, viz., the presence of flagella—not 
postical but lateral branches, attenuated and subaphyllous at the decurved 
and rooting extremity, which brings them into close relationship with 
a group, consisting of Lejewnea auriculata, Wils. (=Phragmicoma versi- 
color, Lehm. et Lg.), Phragmicoma humilis, G. (=Phr. repleta, Tayl.), 
Phr. ligulata, Nees, and a few others, mostly relegated in the ‘Synopsis’ 
to Phragmicoma, but some to Lejeunea. These all agree with typical 
Thysananthus in habit, rooting flagella, lingulate leaves, repeatedly and 
closely innovant inflorescence, and trigonous perianths ; and only differ 
in the entire underleaves and perianths. To this group I have applied 
the name Mastigolejewnea, on account of the flagella, which scarcely exist 
elsewhere in Lejewnea. 
The next genus is Ptychanthus, comprising several species, all Oriental 
(chiefly Malayan) except one South American, Ptych. Theobrome, n. sp., 
found by myself on cacao trees near Guayaquil. Their aspect is almost 
of Bryopteris, but more laxly pinnate, and with more distant leaves. A 
more essential difference resides in the 7—10-plicate perianth, and in the 
branches being at least once—sometimes repeatedly—innovant, and 
floriferous at each successive apex. 
Phragmicoma was founded by Dumortier on a solitary species, Junger- 
mania Mackaw, Hook. As it stands in the ‘Synopsis’ it can only be 
described as an agglomeration of species of various distinct types, one of 
which (Mastigolejewnea) I have indicated above. Of the two sections into 
which the genus is divided in the ‘ Synopsis,’ the first, comprising those 
few species with a perfectly complanate perianth, such as Phr. Mackait, 
Phr. Guilleminiana, and Phr. Bongardiana, is truly a natural group ; but 
one of the characters assigned to it, “ Pedicellus capsule in planta exsic- 
cata levis,” is no character at all; for the pedicel in these species, as in 
all other Lejeunee, is built up of horizontal tiers of cells, which are 
lengthened out at maturity, and the resistance to shrinkage of the lateral 
walls is merely a difference of degree, leaving the dried pedicel nearly or 
quite smooth in several of the robuster species, and nodose at the join- 
ings of the tiers in all the more delicate ones. The definition of “§ 2, 
Ptychanthoides. Perianthium a latere compressum, carina dorsali una 
binisque ventralibus instructum. Pedicellus internodiis (post exsicca- 
tionem) tuberoso-geniculatus ”* is disfigured by two errors ; for the com- 
pression of the perianth is not lateral but frontal, as in every Lejeunea,— 
indeed as in all Jubulee ; and the pedicel in some of the species is as smooth 
as that of Phr. Mackaw. Moreover, the “perianth unicarinate above, 
bicarinate beneath,” along with roundish entire under leaves, exactly corre- 
sponds to a number of species that figure in the ‘Synopsis’ not as 
Phragmicome but as Lejewnew,—such as L. unciloba, Ldng., L. clypeata, 
Schwein., LZ. Leprieurit, Mont., &e. In Phragmicoma is included a 
* ‘Syn. Hep.,’ p. 294, 
