SIJJROTH : ox NEOnXALIMAX. 43 



four bundles, the two shorter and inner ones being inserted in the 

 buccal mass or pharynx, whilst the lateral ones are attached directly 

 to the body-wall on each side of the mouth. These are not pigmented. 

 The right lateral bundle lies on the inner side of the penis. The 

 muscles of the ommatoi^hores, which have slender terminations, unite 

 with these bundles at a point near the cerebral ganglia, and represent 

 only secondary branches of them. Thus the distal ends of the lateral 

 bundles are on the same level as the roots of the ommatophores, 

 the bundles themselves occupying the places of the missing smaller 

 tentacles. 



The pedal gland. — The pedal gland opens below the mouth, and is 

 one-fourth the length of the body. It is sharply circumscribed, and 

 somewhat flocky, the excretory duct being visible from the upper side. 

 It is attached to the body-wall by mescnt(nial tissue. 



Generic fosition. — It is much to be regretted that Fischer, when 

 describing his Uijalimax Maillardl, paid no attention to the separa- 

 tion of the male and female ducts, the tentacles, the columellar 

 muscle, the pedal gland, etc. ; nor have we any information whether 

 the shell envelops any portion of the intestinal sac or not, so 

 that a correct compaiison is very difficult. As far as it is possible 

 to arrive at any conclusion, the Erazilian form agrees perfectly 

 with Hyalimax in its habitat, its mantle, its foot-sole, jaw, radula, 

 and the simple termination of its genital organs, as well as in the lack 

 of accessory glands, dart sac, etc. The sole difference consists in the 

 position of the genital opening. This in the Brazilian form is nearer 

 to the ommatophore than in the true Uijalimax. The difference in the 

 insertion of the receptacular duct is only one of specilic, not of 

 generic, value. It, however, seems very probable, considering the 

 wide geographical separation of the two, that more exact investigation 

 would bring further differences to light. I therefore propose to call 

 the new form Neohyalimax Jirasiliensis, which may rank either as 

 a subgenus of Hyalimax or as a new genus altogether ; the value of 

 the more flattened shell and more forward position of the genital 

 opening being merely subjective. 



Position in the family : (a) Its relation to the Succineidce. — The fore- 

 going description demonstrates that Neohyalimax is closely allied to 

 Succinea, on the basis of the following points of resembh^nce : the 

 elasmognathous jaw; the radula; the early separation of the genital 

 ducts ; the absence of accessory genital glands ; the wide separation of 

 the supra-pharyngeal ganglia ; the fusing together of the pleuro-visceral 

 ganglia ; the position of the heart ; and the absence of foot-sole grooves. 

 Possibly to this may be added the distribution of the blood-vessels in 

 the wall of the pulmonary chamber, the respiratory area being equally 

 divided into an intestinal lung and a columellar lung ("Darmlunge" 

 and " Spindellimge ") in Semper's sense (3). This hypothesis is founded 

 1 on the suggestion that the principal black line in Fig. 4 is identical with 

 the principal pulmonary vein. The relationship of Neohyalimax to 

 the neotropical genus Ilomalonyx, which is placed between Hyalimax 

 and Succinea in the textbooks (Fischer, Pilsbry), is more critical. 

 Decision on this point must be deferred until a further and better 



