Polygala polygama and P. paucifiora. 129 
Chodat,! in speaking of the two winglike sepals in the 
Polygalacee says: “ The anatomical structure of these wings 
is that of the petals. Stomata have never been found on 
them.” The following description will show the incorrectness 
of the above statement. The cells of each surface are wavy 
in outline, similar to the smaller sepals (Fig. 11). On the 
outer surface stomata are numerous and on the inner they are 
also tolerably abundant. In both cases they occur in the 
central portion of the member, but cease abruptly toward the 
margins, where the fibro-vascular bundles end. Thus, taking 
into account the loose and spreading character of the wings, 
it will be seen that over the calyx stomata are found on sur- 
faces which obtain free access of air, and rather irrespective 
of the presence of chlorophyll. 
Calyz.—B. The Aérial Cleistogamic Flower. In this all 
five sepals are present, but the wings are reduced and only 
slightly larger than the other three. The epidermal cells 
show to a less degree the wavy outlines of the corresponding 
cells in the conspicuous flowers (Fig. 12). The superficial 
wall-thickenings are not visible. Onthe outer surfaces of the 
three exterior sepals stomata are exceedingly abundant. 
Calyx.—C. The Subterranean Cleistogamic Flower. In 
this five sepals are still present. The wings are greatly 
reduced and are slightly smaller than the other three sepals. 
The wavy cell-walls, found in the epidermis of the chasmo- 
gamic flower sepals, are here lacking (Fig. 13). Stomata 
are present on all five sepals over both surfaces, though they 
are few in number, and are of a characteristically degenerate 
appearance. 
Corolla——A. The Chasmogamic Flower. In it the ante- 
rior petal is 5-6 millimeters long, and is developed intoa hood, 
covering the sporophylls and bearing on the anterior border of 
the hood six or eight finger-shaped appendages, which pro- 
1Chodat, loc. cit. 
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