134 Shaw—Structure of the Flowers in 
Their rich contents would lend support to this view. Such 
a suggestion has already been made by Mrs. Pettit’ in the 
case of the gynophore hairs of Arachis hypogea. Or since 
such underground fruits would be especially liable to destruc- 
tion from animals living in the soil, these glandular bodies 
may contain some alkaloidal or other substances which may 
protect the fruit against such. The fact that these bodies are 
only found on that portion of the ovarian surface which 
comes into direct contact with the soil as soon as the ovary 
has grown slightly, might favor the former view. Stomata 
have never been found on the subterranean ovary. 
Since the style scarcely exists in the cleistogamic pistil we 
should hardly know what to expect in regard to the passage 
from the ovarian cavity tothe exterior. In point of fact open- 
ings very frequently occur close beside the stigmatic knob. 
Until the flowers are quite well developed (nearly ready for 
pollination), there is always a considerable opening into both 
chambers of the ovary. 
Nectary.—A. In the chasmogamic flower, close under the 
posterior base of the ovary, there is found growing out from 
the receptacle a small tract of cells. These are covered by 
the slightly modified posterior sepal, and receive branches 
from the vascular system of the peduncle. The surface of the 
papilla is formed of irregular polygonal cells. Beneath these 
is a region of cells, elongated perpendicularly to the surface, 
and lying below them in turn are the terminations of the vas- 
cular bundles. There is no doubt as to the character of the 
structure as a nectary. Insects have been observed to alight 
on the summit of the corolla and to probe in this region. 
Nectary. 
of the nectary is marked by faintly developed vascular bun- 
dles, and in some instances bya small cellular papilla. The 
B. In the aérial cleistogamic flower the position 
development of the latter is however very variable. 
1 Pettit, Arachis Hypogea. Memoirs of the Torr. Bot. Club, vol. iv, No. 4. 
