Structure, etc., of Epiphegus Virgintana. 377 
bundles are relatively well developed. There is a core of long 
slender spiral tracheae extending out farthest in the lobes. 
On both sides of this occur short stout spiral tracheids, set 
irregularly and discontinuously along the bundle. A few 
small phloem cells lie to one side. The tissue cells are much 
larger than the bundle cells. They fit together in all kinds 
of positions, except near the bundles, where they arrange 
themselves quite regularly. 
One-celled, rarely two-celled hairs fringe the edges of the 
lobes. Below, across the base of the lobes, there extends 
a band of two or three-celled hairs, longer than the upper 
hairs. All of these hairs are on the outer surface of the 
calyx; none are present on the inner surface. They have a 
swollen granular appearance. 
The Corolla—In the chasmogamic flower (Fig. 3) the 
corolla consists of cells longitudinally elongated, and is of 
four or three cells in thickness. The outer layer of cells 
is quite regular and even, constituting an epidermis. At 
the base, where the corolla passes under the calyx, it 
thickens greatly, being many times thicker than either the 
calyx or the upper corolla region, and consisting of two cell 
layers. Five vascular bundles pass through its length and 
end in the five corolla lobes. They are similar to the bundles 
of the calyx. 
There are two areas of hair development, both on the 
inner surface. One area forms a hairy belt along the outer 
margins of the corolla lobes. These are short, mostly two- 
celled hairs, and often so bend on themselves as to have a 
hooked appearance. They are greatly swollen and turgid- 
looking. The other area is a ring extending round the inner 
surface at about the region where the stamens are inserted. 
These hairs are much longer than those above. They con- 
sist of about three narrow elongated cells. At the free end 
of each hair there is often a flat, flange-like expansion. 
The cleistogamic corolla is greatly reduced in size. It 
becomes an apparently closed asymmetric cap-like structure 
