378 Cooke and Schively on Observations on the 
that is lifted off by the maturing ovary. The bundles are 
also reduced. A small fifth one may appear, but usually 
there are but four bundles. Two of these are fairly large, 
and show both spiral tracheae in three or four rows, and 
tracheids. They occur about one-third of the circumference 
of the corolla apart, and arise on the anterior short curve 
of the cap-like corolla. In that position they can best sup- 
port the corolla. The two small bundles alternate with these 
large ones. They apparently contain but a single spiral 
trachea, that extends clear to the end of the corolla. The 
larger bundles end some distance below. Stomata have 
been found on the corolla. 
This corolla is from four to six cells in thickness. The 
outer layer is of large regular close-set cells, with a con- 
siderable amount of thickening developed on their outer 
walls. A protective epidermis is thus supplied to it. Longi- 
tudinal sections reveal that the corolla is not in strict mor- 
phological sense a cleistogamic one, but has a distinct 
though small mouth (Fig. 4). This opening occurs where 
the anterior straight, or slightly concave and convex sur- 
faces of the corolla meet. The convex surface develops a 
disc of greatly thickened cells that overlaps the concave 
side, leaving a distinct mouth. This mouth is entirely closed 
by the numerous small hairs crowding into it from within 
the corolla. These hairs are of one to two round swollen 
cells. They arise in great numbers on the inner surface 
of the corolla all about this mouth. Also a group of hairs 
appears on the outer surface of the corolla, tip, just where it 
is overlapped by the other surface. These hairs help greatly 
in filling up this mouth. The corolla may then be called 
physiologically but not morphologically cleistogamic. There 
is a slight development of long hairs about the region where 
the stamens arise. 
The hairs closing the corolla’s mouth are evidently in- 
tended to keep out intruding insects. Probably the numer- 
ous hairs developed on the chasmogamic flowers have also a 
