Structure, etc., of Eptphegus Virginiana. 379 
definite purpose with regard to insect visitors. But just 
what this purpose is has not been discovered. 
The Androecitum.—There are four stamens inserted on 
the corolla and one-third from its base. This is true of 
both cleistogamic and evident flowers. But in the cleisto- 
gamic flower the stamens show great reduction in size and 
an actual increase in the production of micro-spores (Fig. 
4). 
In the chasmogamic flower the filament shows a large 
central bundle. At the base, both phloem and xylem are 
well developed. But the phloem constantly decreases in 
amount toward the anther region. Hairs develop across 
the inner face of the filament, just above its origin on the 
corolla. They are nearly identical with the hairs of the 
corresponding area on the corolla, being very long, narrow 
and multicellular. 
The anther consists of two lobes that hang dependent 
from the upper tip of the filament. They are remarkable in 
being so distinctly separated from each other. They meet 
in the upper portion, but diverge widely below, being found 
on opposite sides of the filament. 
There is a great development of hairs even on these anther 
lobes. Each hair is extremely long, slender, unicellular and 
has tapering ends that often hook downward. A great deal 
of thickening is developed in the cells of these anther walls, 
of a clear, shining, refractive appearance. In section, this 
thickening is seen to be deposited quite evenly about the 
walls of the two or three rows of cells that form the anther 
sac. But in the late stage of the anther, this thickening is 
loosened out from the cells and split up in various ways, so 
that bars and angles and even squares of glistening thick- 
ening substance are seen lying over each other in the great- 
est confusion. This state of affairs was always noted when 
dehiscence had occurred, so it was probably a result of 
dehiscence. The line of dehiscence is a vertical one, passing 
from top to bottom of the anther wall. 
