384 Cooke and Schively on Observations on the 
of the capsule, curious as this may seem. The more anterior 
and somewhat right-hand of the two valves always con- 
tains considerably more seed than does the posterior valve, 
950 to 770 being a fairly average ratio. The style is divided 
in two regions, corresponding to these two valves. The pos- 
terior stylar half lies above, next the exterior of the corolla, 
when it has curved over to reach the stamens. So the 
anthers lie adjacent to the lower half of the style, and send 
their pollen tubes in here. Naturally, therefore, more pollen 
tubes pass down the anterior region of the style, and more 
ovules are therefore fertilized in the anterior valve of the 
capsule. 
Fertilization in the chasmogamic flowers is probably by 
means of insects. The nectary and peculiar arrangements 
of the numerous hairs seem to indicate this. Protogyny is 
evidently the rule here. Ona number of plants observed in 
early September, the lower and older evident flowers showed 
a brown, shrunken, dried-up pistil, while the anthers were 
just about mature, some having dehisced, but not yet shed 
their pollen. The upper and younger chasmogamic flowers 
showed a mature pistil, with sticky stigma bearing numerous 
attached pollen grains. The end of the style curves over and 
downward so that insects must brush by it in going to the 
nectary. In these flowers the anthers were decidedly imma- 
ture in appearance. So protogyny is the existing condition. 
THE DEVELOPMENT OF THE OVULE. 
The following observations were made entirely from 
cleistogamic flowers. 
In very young buds, the ovary is seen to be filled with 
pale, faintly staining cells. Through these cells pass vertical 
double rows of very deeply stained cells. These are the 
earliest beginnings of the ovules. The cells are rectangular, 
with dense cytoplasm and a very large nucleus. The nuclear 
cavity is clear and almost unstained. In it are seen great 
