Structure, etc., of Epiphegus Virginiana. 393 
with the beech-root, so to speak. That is, the attachment to 
the beech-root occurs some distance above the base, on the 
side walls of the young Epiphegus. Others form the con- 
nection directly through their base. Evidently the ovules 
giving rise to the latter rested above the beech-root and 
germinated in that position. The seeds must in all proba- 
bility lie beside the beech-root to form the lateral connection. 
Two young tubers attached in the same plane on a beech- 
root, one on either side, have been observed. Mature plants 
sometimes show the same condition. 
In the one here described the connection was lateral. 
Near the point of connection the epidermal cells of Epiphegus 
take on a peculiar darkly colored appearance. At this level 
the several bundle-masses in the tuber seem to stream across 
in nearly parallel direction, converging toward the haustorial 
connection. 
In most cases of parasitism the connection between the 
two plants is formed by the parasite, that pushes its suckers 
into the tissue of the host. But in Epiphegus, all indications 
point to the conclusion that the host sends the “haustorium”’ 
into the parasite (Plate XXX, Fig. 2). The appearance in 
sections through this region is of a number of arms growing 
from the beech-root into the parasite. The peculiar tissue 
composing these arms is totally unlike any tissue found 
either in Fagus or in Epiphegus, and is found only in the 
beech-root at the point of union. The growth of this haus- 
torial organ always occurs at the ends in Epiphegus, not at 
the end in Fagus. And the beech-root remains very small 
and fine (Plate XXIX), even when it is attached to a great 
swollen parasitic tuber of the adult Epiphegus. It certainly 
seems that this organ of connection arises from the beech 
and grows into the tissues of the parasite. It will be assumed 
that this is the case in describing the structure, as it is the 
easiest method for description. 
Near the point of connection the tissues of the beech- 
root assume a thoroughly disorganized appearance. . The 
