July 1891.] BOTANICAL SOCIETY OF EDINBURGH. 159 



The Cotyledonary Glands in some Species of Eubiace^. 

 By Thomas Berwick, St Andrews. 



In a previous paper read to this Society,* I noted the 

 presence of two embryonic glands in tlie axils of the 

 cotyledons of the ungerminated embryos of Galium Aparine, 

 L. ; G, cruciata, Scop. ; and Slierardia arvensis, L. 



Since then I have dissected the ungerminated embryos of 

 numerous other species of the Eubiacete, and find two 

 glands in the axils of the following, viz. : — Asperula 

 arvensis, L. ; A. setosa, Jaub. et Spach. ; Borreria capitellata, 

 Chmss. et Sch. ; Callipdtis Cucullaria, Stev. ; Galium 

 anglicum, Huds. ; G. articulatum, Eoem. et Sch. ; G. 

 horeale, L.- ; G. capillipes, Rchb. ; G. caudatum, Boiss. ; G. 

 lucidum, D.C. ; G. macrocarpum, Boiss. ; G. Mollugo, L. ; G. 

 nehulosum, Boiss. ; G. physocarpum, Boiss. ; G. saccliaratum, 

 All. ; G. spuriiim, L. ; G. tenuissimum, M. B. ; G. iricorne, 

 "Wither. ; Phyllis Nohla, L. ; Spermacoce tenuior, L.; Vaillantia 

 hispida, L. ; V. incrassata, Pomel. ; V. muralis, L. 



The form of these glands in the embryonic stage corre- 

 sponds with that found after germination.+ The two 

 embryonic glands in the plants in the above list keep the 

 lead,t in point of size, of all the other cotyledonary glands 

 which appear during the processes of growth. It would 

 seem from the comparatively large size they attain, when 

 more or less fully developed, that, as is the case in Galium 

 Aparine, they will outmeasure any gland afterwards de- 

 veloped in the leaf whorls of any of the species named. 



In these 26 species of Rubiacese every embryo of the 

 very numerous, and in most cases minute seeds, produces 

 its two isolated glands, varying in their outline and dimen- 

 sions in different species, but fixed in position and identical 

 in form in the same species. 



Multiplication f of cotyledonary glands takes place in 

 almost all the above as was the case in G. Aparine. There 

 is reason to believe, that a larger embryo contains two 

 larger glands than those found in a smaller embryo, but 



* Trans. Bot. Soc. Edin., vol. xviii. p. 436. 

 t Verified in most species in list. 



