382 TRAXSACTIONS AND PROCEEDINGS OF [Sess. lvi. 



possessing a granular border, while the latter seems to be a 

 colloid mass stained by hcematoxylin. In fig. 40^ one large 

 body (?) is partly covered by the nucleolar bag (n. h.). Two 

 minute bodies each with an ill-defined central feebly-stained 

 spot are shown in fig. 41 towards the basal end of the 

 nucleus, while towards its micropylar end a number of 

 spherical bodies, each with a dark central spot, occur, two of 

 which (perhaps a third one on the left side) are outside the 

 nucleus, while two are within the nuclear membrane. Fig. 

 31 evidently shows a primary endosperm-nucleus with a large 

 nucleolus in its upper half and a number of globular bodies, 

 each with a central deeply-stained spot in its lower half. 

 These globular bodies almost seem to have arisen by divi- 

 sion of the large body (?) shown in fig. 40^ Fig. 39 may 

 perhaps throw some light on the manner in which the 

 globular bodies with a dark centre get outside the nuclear 

 membrane, for we see three very distinct spherical bodies 

 lying outside the nucleus at its micropylar end, and a clear 

 strand {k) running from them to the nucleus. A number of 

 indefinite bodies (a; ? and y ?) are seen close to the basal end 

 of the nucleus. How these bodies arise I am unable to say, 

 they may have some connection with the paranucleolar 

 bodies (p. n., figs. 32, 33, 35, 36''), as fig. 35 would make me 

 incline to believe, for here we find two bodies very like para- 

 nucleoli lying in close contact with one another and only 

 separated by the disappearing septum of the niicleus. 



Whether these paranucleoli of two different nuclei have 

 the power of conjugation just as the nucleoli have, I shall 

 endeavour to find out, as soon as I have more time at my 

 disposal ; but a plant with larger nuclei than Myos^trus will 

 have to be taken for examination. 



These bodies outside the nuclear membrane I saw first in 

 September 1890, but was unable to explain them, but now I 

 believe them to Ije identical with the paranuclei of Von la 

 Valette St George, or the " spheres attractives " of v. Beneden, 

 or the " Centrosomata " and " Archoplasm " of Boveri, or 0. 

 Hertwig's " Polkorperchen," * or the " directing bodies " of 

 English authors,t or v. Tieghem's " Tinoleucites." 



* Max Schultze's Archiv. f. mikroscop. Anat. xxxvi., 1890, p. 29. 

 + "Polar bodies" of Euglisli autliors correspond to the " Richtungs- 

 kiJrper " {'i.e., directing bodies) of 0. liertwig. 



