July 1900.] BOTANICAL SOCIETY OF EDINBUKGH 3.") 5 



this I find in P. SchrenJciana (Type 4 staminal arrange- 

 ment) that the staminal papillae of eacli festoon may develop 

 simultaneously. 



Fig. 2 shows a young ilower of this species bisected 

 in the antero-posterior plane. The epicalycine, calycine, 

 and petaliue papilhe are developed, but there is as yet 

 no trace of the staminal papilhe. Points to be noted 

 in this figure are the relatively large size of the epi- 

 calycine papilhe at this early stage — a feature which 

 would in some degree support the view of the morpho- 

 logical nature of the epicalyx suggested in this note. 

 Another interesting point is the close relation of the 

 petaline to the epicalycine papillfe, the former being 

 practically outer growths or branches of the latter. 



By the time that the staminal papillai are all represented 

 the epicalyx and calyx segments are of large size, and, 

 arching inwards, form a funnel-shaped covering over the 

 centre of the fiower : the throat of the funnel is closed 

 by a considerable growth of hairs from the inner surfaces 

 of the segments. These hairs develop very precociously, 

 rudiments of them being found at a very early stage on 

 the papillae. It is only now that the carpellary papilhe 

 begin to appear on the hitherto smooth hemispherical 

 surface of the gynophores. The full details of development 

 of the various types of flowers, and also the discussion 

 as to the nature of the corolla, suggested by its manner 

 of development are reserved for -a future note. 



Each stamen (Fig. 9) is composed of a free filament 

 tapering towards its apex, where it bears a sub-sagittate 

 anther, the lobes of which converge apically and diverge 

 basally. The filament (Fig. 10) has a single central bundle 

 surrounded by a loose cylinder of parenchyma, the cells 

 of which increase in size, progressing outwards to tlie 

 epidermis. The anther lobes are two in number in the 

 immature stamens (Fig. 11), and the dehiscence is longi- 

 tudinal and sub-lateral. The anther wall consists of two 

 layers, an inner fibrous and an epidermis. Where the 

 epidermis dips into the connective between the anther lobes 

 on the posterior and anterior faces, its cells are much 

 elongated radially, and contain a pigment which stains 

 deeply with htematoxylin (Fig. 11,^;.). 



