338 TKANSACnONS AND Pl:OCEEDI^'GS OF Till': [i5i.^s. i.xu. 



to sub-anatropous conditions when the raphe is hardly 

 marked. 



The Ovarian ITall is usually white in colour, but becomes 

 brown on maturing. The whole ovary also increases some- 

 what in size, and the style drops off on fruiting. The 

 ovaries are usually closely packed together on the torus, 

 and entirely concealed by the projecting basal epidermal 

 flanges of the styles. 



The ovarian wall (Fig. 3) is composed of three strata of 

 tissue — (1) an outer parenchymatous (p.) of a few layers of 

 closely packed cells, the epidermal having cutinised walls ; 

 (2) a middle crystallogenous layer {<!x.) of small, close set,. 

 quadrangular, thickish-walled cells, each with a cubical 

 crystal of oxalate of lime ; (3) an inner sclerenchymatous 

 stratum composed of {a) an outer layer of spindle-shaped 

 fibres parallel to the long axis of the ovary, and (b) an 

 inner layer of similar fibres with their long axes transverse 

 to that of the ovary. 



The Sti/le is continuous with the ventral keel of the 

 ovary, to which it is fixed by a very constricted neck. 

 The length of the style varies, and this may be a feature 

 of systematic importance, as there are well marked long and 

 short styled species. It is composed (Fig. 7) of a cylinder 

 of parenchyma filled with a loose conducting tissue of cells, 

 rich in proteids. The parenchyma cells increase in size 

 outwards to the epidermis, which is composed of larger 

 cells protruding outwards, often in a papillose fashion. 

 These cells are specially noticeable just at the base of the 

 style (Figs. 8 and 14, h.f.) near tlie constricted neck, where 

 they form a sort of basal flange or rosette. They are less well' 

 developed on the ventral side, hence appears a ventral 

 groove, which is continuous with a gap in the lip of the 

 funnel-shaped papillose stigma (FMgs. 15 and 16). There is 

 usually a more or less well markeil constriction just beneath 

 stigma. 



All the outer cells of the style contain a peculiar 

 pigment of a light lemon -yellow colour, apparently 

 similar to that already mentioned as occurring in the 

 connective epidermis of the anthers. It resists the 

 usual solvents, gives a brownish precipitate with hydro- 

 chloric acid, and dissolves, after a ])rcliininary change 



