122 TRANSACTIONS OF THE [Sess. lxxxi 



on the outside (fig. 8). The stamens have the typical apical 

 appendage and short, obtuse, basal appendages, the con- 

 tiguous appendages being more or less connate (figs. 4 and 

 5). The corolla is tubular, regularly and deeply 5-lobed ; 

 the style is not exerted (fig. 2). 



The structure of the style and stamens and the presence 

 of filiform female florets at once suggests the Inuleae. 

 According to Bentham's classification of the order the new 

 genus falls into the sub-tribe Plucheineae of the Inuleae, 

 and from the floral characters should be placed near 

 Pluchea, from which it is distinguished by habit, receptacle, 

 and inflorescence. A few species of Phicliea are herbaceous 

 perennials, but most are shrubby. The capitulum in 

 Pluchea is usually small and the inflorescence corymbose, 

 but at least one species (P. aroiinatica, Balf. f. from Socotra) 

 shows large capitula and a diflusely corymbose inflores- 

 cence. The receptacle is naked and the anther tails are 

 connate and acuminate in Pluchea. The new genus is 

 distinguished from BLumea and Laggera by the undivided 

 style of the male florets, fimbrillate receptacle, general 

 habit, and quite a few other characters ; and no other genus 

 in the Inuleae approaches it closely. 



The large percentage of capitula with no male florets is 

 interesting as showing a tendency to dioecism, but the most 

 interesting point is the placing of the plant in Saussurea 

 by Druminond. In Table I of a paper ^ by one of us the 

 Inuleae are shown to be more closely allied in their anther 

 appendages to the Mutisieae and Gynareae than to the 

 tribes among which tliey are usually placed. The typical 

 style of the Inuleae closely approaches some of the Muti- 

 sieae and exceptional Gynareae. From the study of all 

 factors, including geographical distribution, it seems probable 

 that the Inuleae gave rise to the Gynareae in the eastern 

 part of the Mediterranean region, through the Buphthal- 

 meae, so that it is not surprising that, in the absence of an 

 investigation of the filiform florets, this plant should have 

 been classed in the Gynareae. The absence of the ring of 

 hairs on the style and the character of the anther ap- 

 pendages, however, would, even then, place it nearer the 



1 Small, J., 'I'he Polleii-jiresentation Mechaiii.siii in Ihe Coinpositae. 

 Anriai.s of Botany, vol. xxix, No. cxv(1915), p. 457. 



