436 APPENDIX. No. V. 
this cohesion is partial, and such as I have now described, being lodged in the 
tubular interstices ; their pomts extending to the base of the ovarium. From 
these sheaths, to which they are exactly adapted, the antherze seem to be dis- 
engaged in consequence of the unequal growth of the different parts of the 
filament ; the inflected portion ceasing to increase in length at an early period, 
while that below the curvature continues to elongate considerably until the 
extrication is complete, when expansion takes place. 
It is singular that this mode of cohesion between the ovarium and calyx in 
certain genera of Melastomaceze, and the equally remarkable zstivation of 
anther accompanying it, should have been universally overlooked, especially in 
the late monograph of M. Bonpland ; as both the structure and economy cer- 
tainly exist in some, and probably in the greater part, of the plants which that 
author has figured and described as belonging to Rhexia. 
On the limits, structure, and generic division of Melastomaceze, I may 
remark, 
Ist. That Memecylon, as M. du Petit Thouars has already suggested,* and 
Petaloma of Swartz+ both belong to this order, and connect it with Myrtacee, 
from which they are to be distinguished only by the absence of the pelluad 
glands of the leaves and of other parts, existing in all the genera really 
belonging to that extensive family. 
2dly. There are very few Melastomaceze in which the ovarium does not in 
some degree cohere with the tube of the calyx; Meriana, properly so called, 
being, perhaps, the only exception. 
And in the greater number of instances where, though the ovarium is 
coherent, the fruit is distinct, it becomes so from the laceration of the connect- 
ing processes already described. 
3dly. That the generic divisions of the whole order remain to be established. 
On examination, I believe, it will be found that the original species of the 
Linnean genera, Melastoma and Rhewia, possess generic characters sufficiently 
distinguishing them from the greater part of the plants that have been since 
added to them by various authors. In consequence of these additions, how- 
ever, their botanical history has been so far neglected, that probably no genuine 
species of Melastoma, and certainly none of Rhexia, has yet been published in 
M. Bonpland’s splendid and valuable monographs of these two genera. 
* Mélanges de Botanique ; Observ. address. a M. Lamarck, p. 57, 
+ Flor. Ind. Occid. 2, p. 831, tab. 14. 
