232 
Às regards the morphological value of the cupule, whether 
it has to be considered a sporophyll or a modified pinnule 
of a compound leaf, or an integument, | will discuss that 
later on. We may now consider the place where the seed 
was attached to the plant bearing it. Scott says about 
this (133). , We have seen that the structure of the pedicel 
shows that the seed-bearing organ formed part of a leaf. 
Evidence from closely allied species leaves no doubt that 
the seeds were borne on compound fronds or pinnae, 
only differing from the sterile foliage in the suppression 
of the laminae of the leaflets, or rather, perhaps, their 
modification to form cupules” (p. 393). 
Characteristic are the great prominent multicelled glan- 
dular capitate hairs, which are found on the pedicel and 
cupule. 
Within the cupule stands the seed properly, composed 
of a nucellus, enveloped by an integument. This integu- 
ment or testa is concrescent with the nucellus to about 
four-fifths of its height. Round the apex of the nucellus 
it is free and forms the so-called canopy. In this region 
the integument is segmented by septa in about nine loculi 
or chambers, whilst lower, these septa disappear and 
make place for deep furrows in the inner surface. Grad- 
nally these furrows dy out to the place of insertion of 
the canopy. The loculi of the canopy are filled with a 
delicate tissue, which is generally not preserved, and a 
single tracheal strand in each chamber. 
Surrounding the micropyle small ridges can be seen, but 
as they overlie the well-marked septa between the cham- 
bers of the canopy, it is quite possible, as Oliver (104) 
remarks, that ,they owe their prominence to post-mortem 
contraction or collapse of the filling-tissues of the cham- 
bers”’ (p. 105) and thus should not be mistaken for the 
unitlobes of the integument. 
The outer side of the testa is formed by a closed layer 
