239 
is formed by cells which have a horizontal orientation. 
It is perhaps possible that this outer zone represents the 
terminations of the longitudinal fibres. At any rate there 
is little doubt about these two zones being homogen 
in origin. 
Though not preserved in a very good condition there 
seems to have been a thin layer of delicate tissue at the 
innerside of the sclerotesta, the endotesta, but when 
present, it is only to be seen in the micropylar region. 
The stalk of the seed consists of sclerotic tissue in the 
centre, surrounded by parenchyma, probably these layers 
are continued into the sclerotesta and sarcotesta of the 
body of the seed. 
The nucellus has always been found standing free within 
the integument, a large space being left between nucellus 
and sclerotesta (perhaps endotesta) from the chalazal region. 
Within the nucellus is a large megaspore. The pollen- 
chamber has a bell-form with a narrow beak at its top 
pointing into the micropylar tube. 
The vascular organisation consists of a sarcotestal system 
without, and a nucellar one within. The main vascular 
strand enters the chalaza and lets off several branches 
into the sarcotesta, running opposite the secondary ridges 
of the sclerotesta. However, they do not run opposite 
all the secondary ridges, only opposite the outer ones of 
each complex of three, never opposite the middle one. 
Their total number therefore seems to be six. 
The nucellar system is formed by the continuation of 
the chalazal strand, which entering the nucellus, spreads 
out in its lower part, splitting up there into a number of 
bundles, which can be traced through the whole length 
of the nucellus to the base of the pollenchamber. 
