248 
The testa becomes differentiated into the three layers, 
the middle one becoming sclerosed in a later age. In the 
ripe ovule the testa projects as a thick margin round the 
narrow external opening of the micropyle. 
»As regards position the ovule of Sfangeria clearly 
corresponds to a microsporangium or to a sorus of micro- 
sporangia”” (p. 301). The testa of the ovule would be 
homologous with the sporangial wall of the microsporan- 
gium and the free portion of the testa above the nucellus, 
unrepresented in the microsporangium: ,,an annularupgrowth, 
around the apex of the nucellus, of the bulky sporangial 
wall, or which comes to the same thing, of the edge of 
the receptacle, which had kept pace with the single 
sporangium' (p. 302). 
»We compare the ovule to a sorus consisting of a 
single sporangium, which develops in the whole similarly 
to a microsporangium, save that it is bulkier, and the 
wall from the first thicker”” (p. 302). 
In the discussion I will say more about this meaning. 
Now I shall mention a case in which Stopes (146) has 
found lobes in Sfangeria schizodon, at the base of the 
ovule outside the testa which in some cases became so 
large ,that they unite to completely enclose and cover 
over the growing ovule, even after it has attained a con- 
siderable size.” ,, The outgrowth I observed in Sfangeria 
is entirely free from the outer integument of the ovule, 
it is undifferentiated in character, and its tissues are those 
of the sporophyll. It is obviously not split off from the 
outer integument, but is the free upgrowth round the 
already doubly integumented ovule’” (146 p. 565). 
Às ÎI lateron will discuss Stopes regards the sarcotesta 
of the Cycadean seeds to be homologous with the cupule 
of Lagenostoma and therefore she calls them double-inte- 
gumented ones. 
